See also:blood away from the heart to the tissues . As, after
See also:death, they are always found empty, the older anatomists believed that they contained air, and to this belief they owe the name, which was originally given to the
See also:windpipe (trachea) . Two
See also:great trunks, the aorta and pulmonary artery, leave the heart and
See also:divide again and again until they become minute vessels to which the name of arterioles is given . The larger trunks are fairly
See also:constant in position and receive definite names, but as the smaller branches are reached there is an increasing inconstancy in their position, and anatomists are still undecided as to the normal, i.e most frequent, arrangement of many of the smaller
See also:arteries . From a
See also:common-sense point of view it is probably of greater importance to realize how variable the distribution of small arteries is than to remember the names of twigs which are of neither surgical nor morphological importance . Arteries adapt themselves more quickly than most other structures to any
See also:mechanical obstruction, and many of the differences between the arterial systems of Man and other animals are due to the
See also:assumption of the erect position . Many arteries are tortuous, especially when they supply movable parts such as the
See also:face or
See also:scalp, but when one or two
See also:sharp bends are found they are generally due to the artery going out of its way to give off a constant and important branch . Small arteries unite or anastomose with others near them very freely, so that when even a large artery is obliterated a
See also:collateral circulation is carried on by the rapid increase in
See also:size of the communications between the branches coming off above and below the point of obstruction . Some branches, however, such as those going to the basal ganglia of the
See also:brain and to the
See also:spleen, are known as "end arteries," and these do not anastomose with their neighbours at all; thus, if one is blocked, arterial blood is cut off from its
See also:area of supply . As a
See also:rule, there is little arterial
See also:anastomosis across the
See also:line of the
See also:body near the
See also:surface, though the scalp, lips and
See also:thyroid body are exceptions . The distribution of the pulmonary artery is considered in connexion with the anatomy of the lungs (see
See also:SYSTEM) . That of the aorta will now be briefly described .
The Aorta lies in the cavities of thethorax and
See also:abdomen, andARTERIES arises from the
See also:base of the
See also:left ventricle of the heart . It ascends forward, upward, and to the right as far as the level of Aorta. the second right costal
See also:cartilage, then runs backward, and to the left to reach the left side of the body of the 4th thoracic vertebra, and then descends almost vertically . It thus forms the arch of the aorta, which
See also:arches over the
See also:root of the left
See also:lung, and which has attached to its
See also:concave surface a fibrous
See also:cord, known as the obliterated ductus arteriosus, which connects it with the left branch of the pulmonary artery . The aorta continues its course downward in close relation to the bodies of the thoracic vertebrae, then passes through an opening in the diaphragm (q.v.), enters the abdomen, and descends in front of the bodies of the lumbar vertebrae as low as the 4th, where it usually divides into two terminal branches, the common iliac arteries . Above and behind the
See also:angle of bifurcation, however, a long slender artery, called the middle sacral, is prolonged downward in front of the sacrum to the end of the coccyx . It will be convenient to describe the distribution of the arteries under the following headings: (I) Branches for the
See also:neck and upper limbs; (2) branches for the viscera of the thorax and abdomen; (3) branches for the walls of the thorax and abdomen; (4) branches for the pelvis and
See also:lower limbs . The branches for the head, neck and upper limbs arise as three large arteries from the transverse
See also:part of the aorta; they are named innominate,"left common
See also:carotid and left subclavian . The innominate artery is the largest and passes upward and to the right, to the root of the neck, where it- divides into the right common carotid and the right subclavian . The carotid arteries supply the two sides of the head and neck; the subclavian arteries the two upper extremities . The common carotid artery runs up the neck by the side of the windpipe, and on a level with the upper border of the Carotid thyroid cartilage divides into the
See also:internal and
See also:external system. carotid arteries . The internal carotid artery ascends through the carotid canal in the temporal
See also:bone into the
See also:cranial cavity . It gives off an ophthalmic branch to the eyeball and other contents of the orbit, and then divides into the anterior and middle cerebral arteries .
The middle cerebral artery extends outward into the Sylvian fissure of the brain, and supplies the
See also:island of Reil, the orbital part, and the
See also:outer face of the frontal
See also:lobe, the parietal lobe, and the temporo-sphenoidal lobe; it also gives a choroid branch to the choroid plexus of the velum interpositum . The anterior cerebral artery supplies the inner face of the hemisphere from the anterior end of the frontal lobe as far back as the internal parieto-occipital fissure . At the base of the brain not only do the two internal carotids anastomose with each other through the anterior communicating artery, which passes between their anterior cerebral branches, but the internal carotid on each side anastomoses with the posterior cerebral branch of the basilar, by a posterior communicating artery . In this manner a vascular circle, the circle of Willis, is formed, which permits of freedom of the arterial circulation by the anastomoses between arteries not only on the same side, but on opposite sides of the mesial
See also:plane . The vertebral and internal carotid arteries, which are the arteries of supply for the brain, are distinguished by lying at some
See also:depth from the surface in their course to the
See also:organ, by having curves or twists in their course, and by the
See also:absence of large collateral branches . The external carotid artery ascends through the upper part of the side of the neck, and behind the lower
See also:jaw into the parotid gland, where it divides into the internal maxillary and superficial temporal branches . This artery gives off the following branches :—(a)
See also:Superior thyroid to the larynx and thyroid body; (b) Lingual to the
See also:tongue and sublingual gland; (c) Facial to the face, palate, tonsil and sub-maxillary gland; (d) Occipital to the sterno-mastoid muscle and back of the scalp; (e) Posterior auricular to the back of the ear and the adjacent part of the scalp; (f) Superficial temporal to the scalp in front of the ear, and by its transverse facial branch to the back part of the face; (g) Internal maxillary, giving
See also:muscular branches to the muscles of mastication, meningeal branches to the dura mater, dental branches to the teeth, and other branches to the
See also:nose, palate and tympanum; (h) Ascending pharyngeal, which gives branches to the pharynx, palate, tonsils and Jura mater . The subclavian artery is the commencement of the great arterial trunk for the upper
See also:limb . It passes across the root of the neck and behind the clavicle, where it enters the armpit, and Sub-becomes the axillary artery; by that name it extends ciavian as far as the posterior
See also:fold of the axilla, where it enters system. the upper
See also:arm, takes the name of brachial, and courses as far as the
See also:bend of the
See also:elbow; here it bifurcates into the radial and ulnae arteries . From the subclavian part of the trunk the following branches arise :—(a) Vertebral, which enters the foramen at the root of the transverse
See also:process of the 6th cervical vertebra, ascends through the corresponding foramina in the vertebrae above, lies in a groove on the arch of the
See also:atlas, and enters the
See also:skull through the foramen magnum, where it joins its
See also:fellow to
See also:form the basilar artery; it gives off muscular branches to the deep muscles of the neck,
See also:spinal branches to the spinal cord, meningeal branches to the dura mater. and an inferior cerebellar branch to the under surface of the cerebellum . The basilar artery, formed by the junction of the two vertebrals, extends from the lower to the upper border of the pons Varolii; it gives off transverse branches to the pons, auditory branches to the internal ear, inferior cerebellar branches to the under surface of the cerebellum, whilst it breaks up into four terminal branches, viz. two superior cerebellar to the upper surface of the cerebellum, and two posterior cerebral which supply the tentorial and mesial aspects of the temporo-sphenoidal lobes, the occipital lobes, and the posterior convolutions of the parietal lobes . (b) Thyroid
See also:axis, which immediately divides into the inferior thyroid, the supra-scapular, and the transverse cervical branches; the inferior thyroid supplies the thyroid body, and gives off an ascending cervical branch to the muscles of the neck; the supra-scapular supplies the muscles on the dorsum scapulae; the transverse cervical supplies the trapezius and the muscles attached to the vertebral border of the scapula .
(c) Internal mammary supplies the anterior surface of the walls of thechest and abdomen, and the upper surface of the diaphragm . (d) Superior intercostal supplies the first intercostal space, and by its deep cervical branch the deep muscles of the back of the neck . The axillary artery supplies thoracic branches to the
See also:wall of the chest, the
See also:pectoral muscles, and the
See also:fat and glands of the axilla; an acromio-thoracic to the parts about the acromion; anterior and posterior circumflex branches to the
See also:shoulder joint and deltoid muscle; a subscapular branch to the muscles of the posterior fold of the axilla . The brachial artery supplies muscular branches to the muscles of the upper arm; a nutrient branch to the humerus; superior and inferior profunda branches and an anastomotic to the muscles of the upper arm and the region of the elbow joint . The ulnas artery extends down the ulnar side of the front of the fore-arm to the palm of the
See also:hand, where it curves outward toward the thumb, and anastomoses with the superficial volar or other branch of the radial artery to form the superficial palmar arch . In the fore-arm the ulnar gives off the interosseous arteries, which supply the muscles of the fore-arm and give nutrient branches to the bones; two recurrent branches to the region of the elbow; carpal branches to the
See also:wrist joint: in the hand it gives a deep branch to the deep muscles of the hand, and from the superficial arch arise digital branches to the sides of the little,
See also:ring, and middle fingers, and the ulnar border of the
See also:finger . The radial artery extends down the radial side of the front of the fore-arm, turns
See also:round the outer side of the wrist to the back of the hand, passes between the 1st and 2nd metacarpal bones to the palm, where it joins the deep branch of the ulnar, and forms the deep palmar arch . In the fore-arm it gives off a recurrent branch to the elbow joint; carpal branches to the wrist joint; and muscular branches, one of which, named superficial volar, supplies the muscle of the thumb and joins the ulnar artery: in the hand it gives off a branch to the thumb, and one to the radial side of the index, interosseous branches to the interosseous muscles, perforating branches to the back of the hand, and recurrent branches to the wrist . The branches of the aorta which supply the viscera of the thorax are the coronary, the oesophageal, the bronchial and the pericardiac . Visceral The coronary arteries, two in number, are the first branches Viasceral of the aorta, and arise opposite the anterior and left posterior segments of the semilunar
See also:valve, from the wall of the aorta, where it dilates into the sinuses of Valsalva . They supply the tissue of the heart . The oesophageal, bronchial and pericardiac branches are sufficiently described by their names .
The branches of the aorta which supply the viscera of the abdomen arise either singly or in pairs . The single arteries are the coeliac axis, the superior mesenteric, and the inferior mesenteric, which arise from the front of the aorta; the pairs are the capsular, the two renal, and the two spermatic or ovarian, which arise from its sides . The single arteries supply viscera which are either completely or almost completely invested by the peritoneum, and the
See also:veins corresponding to them are the roots of the vena portae . The pairs of arteries supply viscera
See also:developed behind the peritoneum, and the veins corresponding to them are rootlets of the inferior vena cava . The coeliac axis is a thick,
See also:short artery, which almost immediately divides into the gastric, hepatic and splenic branches . The gastric gives off oesophageal branches and then runs along the lesser curvature of the stomach . The hepatic artery ends in the substance of the
See also:liver; but gives off a cystic branch to the
See also:gall bladder, a pyloric branch to the stomach, a gastro-duodenal branch, which divides into a superior pancreatico-duodenal for the pancreas and duodenum, and a right gastro-epiploic for the stomach and omentum . The splenic artery ends in the substance of the spleen; but gives off pancreatic branches to the pancreas,
See also:vasa brevia to the left end of the stomach, and a left gastro-epiploic to the stomach and omentum . The superior mesenteric artery gives off an inferior pancreaticoduodenal branch to the pancreas and duodenum; about twelve intestinal branches to the small intestines, which form in the sub-stance of the mesentery a series of arches before they end in the wall of the intestines; an ileocolic branch to the end of the ileum, the caecum, and beginning of the colon; a right colic branch to the ascending colon; and a middle colic branch to the transverse colon . The inferior mesenteric artery gives off a left colic branch to the descending colon, a sigmoid branch to the iliac and pelvic colon, and ends in the superior haemorrhoidal artery, which supplies the rectum . The arteries which supply the coats of the alimentary
See also:tube from the oesophagus to the rectum anastomose freely witheach other in the wall of the tube, or in its mesenteric
See also:attachment, and the anastomoses are usually by the formation of arches or loops between adjacent branches . The capsular arteries, small in size, run outward from the aorta to end in the supra-renal capsules .
The renal arteries pass one to each
See also:kidney, in which they for the most part end, but in the substance of the organ they give off small perforating branches, which
See also:pierce the capsule of the kidney, and are distributed in the surrounding fat . Additional renal arteries are fairly common . The spermatic arteries are two long slender arteries, which descend, one in each spermatic cord, into the scrotum to supply the testicle . The corresponding ovarian arteries in the
See also:female do not leave the abdomen . The branches of the aorta which supply the walls of Parietal the thorax, abdomen and pelvis, are the intercostal, the branches lumbar, the phrenic, and the middle sacral . The intercostal arteries arise from the back of the thoracic aorta, and are usually nine pairs . They run round the sides of the vertebral bodies as far as the commencement of the inter-costal spaces, where each divides into a dorsal and a proper intercostal branch; the dorsal branch passes to the back of the thorax to supply the deep muscles of the spine; the proper intercostal branch (AB.) runs outward in the intercostal space to supply its muscles, and the lower pairs of intercostals also give branches to the diaphragm and wall of the abdomen . Below the last
See also:rib a subcostal artery runs . The lumbar arteries arise from the back of the abdominal aorta, and FIG. i.—Diagram of a pair are usually four pairs . They run of intercostal arteries. round the sides of the lumbar verte- Ao, The aorta transversely brae, and divide into a dorsal branch divided, giving off at which supplies the deep muscles of each side an inter- the back of the loins, and an abdominal costal artery . branch which runs outward to supply PB, The posterior or dorsal the wall of the abdomen . The dis- branch .
tribution of the lumbar and inter- AB, The anterior or proper costal arteries exhibits a trans- intercostal branch. versely segmented arrangement of
See also:IM, A transverse section the vascular system, like the trans- through the internal versely segmented arrangement of mammary artery. the bones, muscles and nerves met with in these localities, but more especially in the thoracic region . The phrenic arteries, two in number, pass to supply the under surface of the diaphragm . The middle sacral artery, as it runs down the front of the sacrum, gives branches to the back of the pelvic wall . Injections made by
See also:Sir W .
See also:Turner have shown that, both in the thoracic and abdominal cavities, slender anastomosing communications exist between the visceral and parietal branches . The arteries to the pelvis and
See also:hind limbs begin at the bifurcation of the aorta into the two common iliacs . The common iliac artery, after a short course, divides into the internal and external iliac arteries . The internal iliac enters the pelvis and divides into branches for the supply of the pelvic walls Iliac and viscera, including the
See also:organs of generation, and for the tem . great muscles of the buttock . The external iliac descends system. behind Poupart's ligament into the thigh, where it takes the name of femoral artery . The femoral descends along the front and inner surface of the thigh, gives off a profunda or deep branch, which, by its circumflex and perforating branches, supplies the numerous muscles of the thigh; most of these extend to the back of the limb to carry blood to the muscles situated there . The femoral artery then runs to the back of the limb in the
See also:ham, where it is called popliteal artery .
The popliteal divides into two branches, of which one, called anterior tibial, passes between the bones to the front of the
See also:leg, and then downward to the upper surface of the
See also:foot; the other, posterior tibial, continues down the back of the leg to the
See also:sole of the foot, and divides into the internal and external plantar arteries; branches proceed from the external plantar artery to the sides of the toes, and constitute the digital arteries . From the large arterial trunks in the leg many branches proceed, to carry blood to the different structures in the limb . The wall of an artery consists of several coats (see fig . 2) . The outermost is the tunica adventitia, composed of connective tissue; immediately internal to this is the yellow elastic coat; Structure within this again the muscular coat, formed of involuntary 01 muscular tissue, the contractile fibre-cells of which are arteries. for the most part arranged transversely to the long axis of the artery; in the larger arteries the elastic coat is much thicker than the muscular, but in the smaller the muscular coat is relatively strong; the vaso-motor nerves terminate in the muscular coat . In the first part of the aorta, pulmonary artery and arteries of the retina there is no muscular coat . Internal to the muscular coat is the elastic fenestrated coat, formed of a smooth elastic membrane perforated by small apertures . Most internal of all is a layer of endothelial cells, which form the
See also:free surface over which the blood flows . The arteries are not nourished by the blood which flows through them, but by minute vessels, vasa vasorum, distributed in their external, elastic and muscular coats .
See also:EMBRYOLOGY The earliest appearance of the blood vessels is dealt with under VASCULAR SYSTEM . Here will be briefly described the
See also:fate of the
See also:main vessel which carries the blood away from the truncus arteriosus of the developing heart (q.v.) . This ventral aorta, if traced forward, soon divides into two lateral parts, the explanation being that there were originally two vessels, side by side, which fused to form the heart, but continued sepa-
See also:rate anteriorly .
The two parts run for a little i distance toward the head oc. of the embryo, ventral to the alimentary canal, and then turn toward the Lv. dorsum, passing one on either side of that tube to 4 form the first aortic arch . Having reached the dor- sum they turn backward toward the tail end and 6 i~vti form the dorsal aortae; here, according to A . H .
See also:Young (Studies in Anatomy,
See also:College, 1891 and 1900) they again turn toward the ventral side and become, after a transitional stage, the hypo-gastric, placental, allantoic or umbilical arteries . This authority does not believe that the middle sacral artery of the adult is the real continuation of the single median dorsal aorta into which the two parallel dorsal vessels just mentioned soon coalesce, though until recently it has always been so regarded . The anterior
See also:loop between the ventral and dorsal aortae already de-scribed as the first aortic arch is included in the maxillary or first visceral arch of the soft parts (see fig . 3, 1) . Later, four other well-marked aortic arches grow behind this in the more caudal visceral arches, so that there are altogether five arterial arches on each side of the pharynx, through which the blood can pass from the ventral to the dorsal aorta . Of these arches the first soon disappears, but is probably partly represented in the adult by the internal maxillary artery, one branch of which, the infraorbital, is enclosed in the upper jaw, while another, the inferior dental, is surrounded by the lower jaw . Possibly the ophthalmic artery also belongs to this arch . The second arch also disappears, but the posterior auricular and occipital arteries probably
See also:spring from it, and at an early
See also:period it passed through the stapes as the transitory stapedial artery . The third arch forms the beginning of the internal carotid .
See also:fourth arch becomes the arch of the adult aorta, between the origins of the left carotid and left subclavian, on the left side, and the first part of the right subclavian artery on the right . The apparent fifth arch on the left side (fig . 3, 6) remains all through foetal
See also:life as the ductus arteriosus, and, as the lungs develop, the pulmonary arteries are derived from it . J . E . V . Boas and W .
See also:Zimmermann have shown that this arch is in reality the
See also:sixth, and that there is a very transitory true fifth arch in front of it (fig . 3, 6) . The part of the ventral aorta from which this last arch rises is a single median vessel due to the same
See also:fusion of the two
See also:primitive ventral aortae which precedes the formation of the heart, but a
See also:spiral septum has appeared in it which divides it in such a way that while the anterior or cephalic arches communicate with the left ventricle of the heart, the last one com- municates with the right (see HEART) . The fate of the ventral and dorsal
See also:longitudinal vessels must now be followed . The fused part of the two ventral aortae, just in front of the heart, forms the ascending part of the adult aortic arch, and where this trunk divides between the fifth and fourth arches (strictly speaking, the sixth and fifth), the right one forms the innominate (fig .
3, In.) and the left one a very short part of the transverse arch of the aorta until the fourth arch comes off (see fig . 4) . From this point to the origin of the third arch is common carotid, and after that, to the head, external carotid on each side . The dorsal longitudinal arteries on the head side of the junction with the third arch form the internal carotids . Between the third and fourth arches they are obliterated, while on the caudal side of this, until the point of fusion is reached on the dorsal side of the heart, the left artery forms the upper part of the dorsal aorta while the right entirely disappears . Below this point the thoracic and abdominal aortae are formed by the two primitive dorsal aortae which have fused to form a single median vessel . As the limbs are developed, vessels bud out in them . The subclavian for the arm comes from the fourth aortic arch on each side, while in the leg the main artery is a branch of the caudal arch which is curving ventralward to form the umbilical artery . From the convexity of this arch the internal iliac and sciatic at first carry the blood to the limb, as they do permanently in
See also:reptiles, but later the external iliac and femoral become developed, and, as they are on the concave side of the bend of the
See also:hip, while the sciatic is on the
See also:convex, they have a mechanical
See also:advantage and become the permanent main channel . . Fin further details see O . Hertwig, Handbuch der vergleichenden and experimentellen Entwickelungslehre der Wirbeltiere (
See also:Jena, 1905') .
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