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CETACEA (from the Gr. ,Eros, a whale)

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Originally appearing in Volume V05, Page 775 of the 1911 Encyclopedia Britannica.
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CETACEA (from the Gr. ,See also:Eros, a See also:whale)  , the name of the mammalian See also:order represented by whales, dolphins, porpoises, &c . From their See also:fish-like See also:form, which is manifestly merely an See also:adaptation to their purely aquatic See also:life, these creatures are often regarded as fishes, although they are true mammals, with warm See also:blood, and suckle their See also:young . by a single crescentic See also:aperture, near the vertex of the See also:head . The bones generally are spongy in texture, the cavities being filled with oil . In the vertebral See also:column, the cervical region is See also:short and immobile, and the vertebrae, always seven in number, are in many See also:species more or less fused together into a solid See also:mass . The odontoid See also:process of the second cervical vertebra, when that See also:bone is See also:free, is usually very obtuse, or even obsolete . In a See also:paper on the form and See also:function of the cervical vertebrae published in the Jenaische Zeitschrift for 1905, Dr O . Reche points out that the shortening and soldering is most pronounced in species which, like the right-whales, live entirely on See also:minute organisms, to See also:capture which there is no See also:necessity to turn the head at all . Accordingly we find that in these whales the whole seven cervical vertebrae are fused into an immovable solid mass, of which the See also:compound elements, with the exception of the first and second, are but little thicker than plates . On the other See also:hand, in the finner-whales, several of which live exclusively on fish, and thus require a certain amount of mobility in the head and See also:neck, we find all the cervical vertebrae much thicker and entirely See also:separate from one another . Among the See also:dolphin See also:group the See also:narwhal and the See also:white See also:whale, or See also:beluga, are distinguished from all other cetaceans by the See also:great See also:comparative length of their cervical vertebrae, all of which are completely free . In the See also:case of the narwhal such an abnormal structure Is easily accounted for, seeing that to use effectively the See also:long tusk with which the male is armed a considerable amount of mobility in the neck is absolutely essential .

The beluga, too, which is believed to feed on large and active fishes, would likewise seem to require mobility in the same region in order to effect their capture . On the other hand, the See also:

porpoise preys on See also:herrings, pilchards and See also:mackerel, which in their densely packed shoals must apparently fall an easy See also:prey with but little exertion on the See also:part of their captor, and we accordingly find all the neck-vertebrae very short, and at least six out of the seven coalesced into a solid Immovable mass . None of the vertebrae are See also:united to form a sacrum . number in mammals, sometimes considerably; they See also:present the exceptional See also:character of having epiphyses at both ends . The See also:pelvis is represented by a pair of small See also:rod-like bones placed longitudinally, suspended below and at some distance from the vertebral column at the commencement of the tail . In some species, to the See also:outer See also:surface of these are fixed other small bones or cartilages, the rudiments of the See also:hind-See also:limb . See also:Teeth are generally present, but exceedingly variable in number . In existing species, they are of See also:simple, See also:uniform character, with conical or compressed crowns and single roots, and are never pre-ceded by See also:milk-teeth . In the See also:whalebone whales teeth are absent (except in the foetal See also:condition), and the See also:palate is provided with numerous transversely placed horny plates, forming the whale-bone." Salivary glands are rudimentary or absent . The See also:stomach is complex, and the See also:intestine simple, and only in some species provided with a small caecum . The See also:liver is little fissured, and there is no See also:gall-See also:bladder . The blood-vascular See also:system is complicated by See also:net-like expansions of both See also:arteries and See also:veins, or retia mirabilia .

The larynx is of See also:

peculiar shape, the See also:arytenoid cartilages and the epiglottis being elongated, and forming a tubular prolongation, The See also:general form is essentially fish-like, the spindle-shaped See also:body passing anteriorly into the head without any distinct neck, and posteriorly tapering gradually towards the extremity of the tail, which is provided with a pair of lateral, pointed expansions of skin supported by fibrous See also:tissue, called " flukes," forming a See also:horizontal triangular propelling See also:organ, notched behind in the See also:middle See also:line . The head is generally large, in some cases attaining more than one-third the entire length; and the mouth is wide, and bounded by stiff, immobile lips . The fore-limbs are reduced to flattened paddles, encased in a continuous skin, showing no See also:external sign of See also:division, and without trace of nails . There are no signs of hind-limbs visible externally . The surface of the skin is smooth and glistening, and devoid of See also:hair, although in many species there are a few bristles in the neighbourhood of the mouth which may persist through life or be present only in the young See also:state . Immediately beneath the skin is a thick layer of See also:fat, held together by a mesh of tissue, constituting the "blubber," which retains the See also:heat of the body . In nearly all species a compressed dorsal fin is present . The See also:eye is small, and not provided with a true lacrymal apparatus . The external See also:ear is a minute aperture in the skin situated at a short distance behind the eye . The nostrils open separately or id a d PMx, Premaxilla . AS, Alisphenoid . Mx, Maxilla .

PS, Presphenoid . ME, Ossified rtion of the See also:

mes- Pt, Pterygoid . ethmoid. pn, Posterior pares . an, Nostrils . Pl, See also:Palatine . Na, Nasal . Vo, Vomer . IP, Inter-parietal . Symphysis of See also:lower See also:jaw . Fr, Frontal. id, Inferior dental See also:canal . Pa, Parietal. cp, Coronoid process of lower jaw . SO, Supra-occipital. cd, Condyle .

ExO, Ex-occipital . See also:

Angle . BO, Basi-occipital. sh, Stylo-hyal . Sq, Squamosal. bh, Basi-hyal . Per, Periotic. th, Thyro-hyal . The lumbar and caudal vertebrae are numerous and large, and, as their See also:arches are not connected by articular processes (zygapophyses), they are capable of free See also:motion in all directions . The caps, or epiphyses, at the end of the vertebral bodies are flattened disks, not uniting until after the See also:animal has attained its full dimensions . There are largely See also:developed See also:chevron-bones on the under See also:side of the tail, the presence of which indicates the distinction between caudal and lumbar vertebrae . In the See also:skull, the See also:brain-case is short, broad and high, almost spherical, in fact (fig . I) . The supra-occipital bone rises upwards and forwards from the foramen magnum, to meet the frontals at the vertex, completely excluding the parietals from the upper region; and the frontals are See also:expanded laterally to form the roof of the orbits . The nasal aperture opens upwards, and has in front of it a more or less horizontally prolonged See also:beak, formed of the maxillae, premaxillae, vomer, and mesethmoid See also:cartilage, extending forwards to form the upper jaw or roof of the mouth .

There are no clavicles . The humerus is freely movable on the scapula at the See also:

shoulder-See also:joint, but beyond this the articulations of the limb are imperfect; the flattened ends of the bones coming in contact, with fibrous tissue interposed, allowing of scarcely any motion . The See also:radius and ulna are distinct, and about equally developed, and much flattened, as are all the bones of the flippers . There are four, or more commonly five, digits, and the number of the phalanges of the second and third always exceeds the normal V . 25which projects into the posterior See also:nares, and when embraced by the soft palate forms a continuous passage between the nostrils and the trachea, or See also:wind-See also:pipe, in a more perfect manner . The brain is relatively large, See also:round in form, with its surface divided into numerous and complex convolutions . The kidneys are deeply lobulated; the testes are abdominal; and there are no vesiculae seminales nor an os penis . The uterus is bicornuate; the See also:placenta non-deciduate and diffuse . The two teats are placed in depressions on each side of the genital aperture . The ducts of the milk-glands are dilated during suckling into large reservoirs, into which the milk collects, and from which it is injected by the See also:action of a muscle into the mouth of the young animal, so that sucking under See also:water is greatly facilitated . Whales and porpoises are found in all seas, and some dolphins and porpoises are inhabitants of the larger See also:rivers of See also:South See also:America and See also:Asia . Their organization necessitates their passing their life entirely in the water, as on See also:land they are absolutely helpless .

They have, however, to rise very frequently to the surface for the purpose of respiration; and, in relation to the upward and downward See also:

movement in the water thus necessitated, the See also:principal See also:instrument of motion, the tail, is expanded horizontally . The position of the nostril on the highest part of the head is important for this mode of life, as it is the only part of the body the exposure II of which above the surface is absolutely necessary . Of numerous erroneous ideas connected with natural See also:history, few are so widespread as that whales spout through their See also:blow-holes water taken in at the mouth . But the " spouting," or " blowing," of whales is nothing more than the See also:ordinary See also:act of expiration, which, taking See also:place at longer intervals than land-animals, is performed with a greater emphasis . The moment the animal rises to the surface it forcibly expels from its lungs the See also:air taken in at the last See also:inspiration, which is charged with vapour in See also:con-sequence of the See also:respiratory changes . This rapidly condensing in the See also:cold See also:atmosphere in which the phenomenon is often observed, forms a column of See also:steam or spray, which has been taken for water . It happens, however, especially when the surface of the ocean is agitated into waves, that the animal commences its expiratory puff before the orifice has cleared the See also:top of the water, some of which may thus be driven upwards with the blast, tending to See also:complete the illusion . From photographs of spouting rorquals, it appears that the height and See also:volume of the " spout " of all the species is much less than was supposed to be the case by the older observers; even that of the huge " See also:sulphur-bottom " (Balaenoptera sibbaldi) averaging only about 14 ft. in height, although it may occasionally reach 20 ft . As regards their See also:powers of See also:hearing, the capacity of cetaceans for receiving (and acting upon) See also:sound-waves is demonstrated by the practice of shouting on the part of the fishermen when engaged in See also:driving a shoal of porpoises or See also:black-fish into shallow water, for the purpose of frightening their intended victims . As regards the See also:possession of a See also:voice by cetaceans, it is stated that one species, the " buckelwal " of the Germans, utters during the breeding-See also:season a prolonged scream, comparable to the scream of a steam-See also:siren, and embracing the whole musical See also:scale, from See also:base to See also:treble . In respect of anatomical considerations, it is true that the external ear is much reduced, the " pinna " being absent, and the See also:tube or " meatus " of very small calibre . On the other hand, the See also:internal auditory See also:organs are developed on the See also:plan of those of ordinary mammals, but display certain peculiar modifications (notably the remarkable See also:shell-like form of the tympanic bone) for intensifying and strengthening the sound-waves as they are received from the water .

It seems, therefore, perfectly evident that whales must hear when in the water . This inference is confirmed by the comparatively small development of the other sense-organs . The eye, for instance, is very small, and can be of little use even at the comparatively small depths to which whales are now believed to descend . Again, the sense of See also:

smell, judging from the rudimentary condition of the olfactory organs, must be in See also:abeyance; and whales have no sense-organs comparable to the lateral-line-system of fishes . Consequently, it would seem that when below the surface of the water they must depend chiefly upon the sense of hearing . Probably this sense is so highly developed as to enable the animals, in the midst of the vibrations made by the See also:screw-like movements of the tail, or flukes, to distinguish the sound (or the vibrations) made by the impact of water against rocks, even in a dead See also:calm, and, in the case of piscivorous species, to recognize by the See also:pulse in the water the presence of a shoal of fish . Failing this explanation, it is difficult to imagine how whales can find their way about in the semi-darkness, and avoid collisions with rocks and See also:rock-See also:bound coasts . In the See also:Christiania Nyt Magazin for Naturvidenskaberne, vol. xxxviii., Dr G . Guldberg has published some observations on the body-temperature of the See also:Cetacea, in which he shows how extremely imperfect is our knowledge of this subject . As he remarks, it is a See also:matter of extreme difficulty to obtain the temperature of living cetaceans, although this has been taken in the case of a white-whale and a dolphin, which some years ago were kept in confinement in a See also:pond in the United States . With the larger whales such a mode of See also:procedure is, however, obviously quite impracticable, and we have, accordingly, to rely on See also:post-mortem observations . The layer of blubber by which all cetaceans are protected from cold renders the post-mortem refrigeration of the blood a much slower process than in most mammals, so that such observations have a much higher value than might at first be supposed to be the case .

Indeed, the blood-temperature of a specimen of See also:

Sibbald's :orqual three days after See also:death still stood at 3 ° C . The various observations that have been taken have afforded the following results in individualcases: Sperm-whale, 40° C.; See also:Greenland right-whale, 38.8° C.; porpoise, 35.6° C.; liver of a second individual, 37.8° C.; See also:common See also:rorqual, 35.4° C.; dolphin, 35.6° C . The See also:average blood-temperature of See also:man is 37° C., and that of other mammals 39° C.; while that of birds is 42 C . The See also:record of 4o° C. in the case of the sperm-whale seems to indicate that' at least some cetaceans have a relatively high temperature . With the possible exception of one See also:West See also:African dolphin, all the Cetacea are predaceous, subsisting on living animal See also:food of some See also:kind . One kind alone (Orca) eats other warm-blooded animals, as See also:seals, and even members of its own order, both large and small . Many feed on fish, others on small floating crustaceans, pteropods and jelly-fishes, while the principal See also:staple of the food of many is constituted by cuttle-fishes and squids . In See also:size cetaceans vary much, some of the smaller dolphins scarcely exceeding 4 ft. in length, while whales are the most See also:colossal of all animals . It is true that many statements of their bulk are exaggerated, but the actual dimensions of the larger species exceed those of all other animals, not even excluding the See also:extinct dinosaurian See also:reptiles . With some exceptions, cetaceans are generally timid, inoffensive animals, active in their movements and affectionate in their disposition towards one another, especially the See also:mother towards the young, of which there is usually but one, or at most two at a See also:time . They are generally gregarious, See also:swimming in herds or " See also:schools," sometimes amounting to many thousands in number; though some species are met with either singly or in pairs . Commercially these animals are of importance on See also:account of the oil yielded by the blubber of all of them; while whalebone, See also:spermaceti and See also:ambergris are still more valuable products yielded by certain species .

Within the last few years whalebone has been sold in America for £2900 per ton, while it is also asserted that £3000 per ton has been paid for two and a See also:

quarter tons at See also:Aberdeen, although there teems to be some degree of doubt attaching to the statement . Soon after the middle of the last See also:century, the See also:price of this commodity was as See also:low as £15o per ton, but, according to Mr See also:Frank See also:Buckland, it suddenly leapt up to £620 with the introduction of " See also:crinoline " into ladies' See also:costume, and it has apparently been on the rise ever since . Ambergris, which is very largely used in See also:perfumery, is solely a product of the sperm-whale, and appears to be a kind of biliary calculus . It generally contains a number of the horny beaks of the cuttlefishes and squids upon which these whales chiefly feed . Its See also:market-price is subject to considerable variation, but from £3 to £4 per oz. is the usual average for samples of See also:good quality . In 1898 a See also:merchant in Mincing See also:Lane was the owner of a lump of ambergris weighing 27o lb, which was sold in See also:Paris for about 85s. per oz., or £18,360 . Whalebone Whales.—Existing Cetacea are divisible into two sections, or suborders, the relationships of which are by no means clearly apparent . The first See also:section is that of the whalebone whales, or Mystacoceti, in which no functional teeth are developed, although there are tooth-germs during foetal life . The palate is furnished with plates of baleen or whalebone; the skull is symmetrical; and the nasal bones form a roof to the nasal passages, which are directed upwards and forwards . The maxilla is produced in front of, but not over, the orbital process of the frontal . The lacrymal is small and distinct from the fugal . The tympanic is welded with the periotic, which is attached to the base of the skull by two strong diverging processes .

The olfactory organ is distinctly developed . The two halves of the lower jaw are arched outwards, their anterior ends See also:

meeting at an angle, and connected by fibrous tissue without any symphysis . All the ribs at their upper extremity articulate only with the transverse processes of the vertebrae; their capitular processes when present not articulating directly with the bodies of the vertebrae . The sternum is composed of a single piece, and articulates only with a single pair of ribs; and there are no ossified sternal ribs . External openings of nostrils distinct from each other, See also:longitudinal . A short conical caecum . When in the foetal state these whales have numerous minute teeth lying in the dental groove of both upper and lower jaws . They are best developed about the middle of foetal life, after which they are absorbed, and no trace of them remains at the time of See also:birth . The whalebone does not make its See also:appearance until after birth; and consists of a See also:series of flattened horny plates, between three and four See also:hundred in number, on each side of the palate, with a See also:bare See also:interval along the middle line . The plates are placed transversely to the long See also:axis of the palate, with short intervals between them . Each See also:plate or blade is somewhat triangular in form, with the base attached to the palate and the See also:apex See also:hanging downwards . The outer edge of the blade is hard and smooth, but the inner edge and apex fray out into long bristly See also:fibres, so that the roof of the whale's mouth ,looks as if covered with hair, as described by See also:Aristotle .

At the inner edge of each principal blade are two or three much smaller or subsidiary See also:

blades . The principal blades are longest near the middle of the series, and gradually diminish towards the front and back of the mouth . The horny plates grow from a fibrous and vascular See also:matrix, which covers the palatal surface of the maxillae, and sends out plate-like processes, one of which penetrates the base of each blade . Moreover, the free edges of these processes are covered with long vascular See also:thread-like papillae, one of which forms the central axis of each of the hair-like fibres mainly composing the blade . A transverse section of fresh whalebone shows that it is made up of See also:numbers of these soft vascular papillae, circular in outline, and surrounded by concentrically arranged epidermic cells, the whole bound together by other epidermic cells, that constitute the smooth (so-called " See also:enamel ") surface of the blade, which, disintegrating at the free edge, allows the individual fibres to become loose and assume a hair-like appearance . Whalebone really consists of modified papillae of the mucous membrane of the mouth, with an excessive and horny See also:epithelial development . The blades are supported and bound together for a certain distance from their base, by a mass of less hardened epithelium, secreted by the surface of the palatal membrane or matrix of the whalebone in the intervals of the plate-like processes . This is the " See also:gum " of the whalers . Whalebone varies much in See also:colour in different species; in some it is almost See also:jet black, in others See also:slate colour, See also:horn colour, yellow, or even creamy-white . In some descriptions the blades are variegated with longitudinal stripes of different hues . It differs also greatly in other respects, being short, thick, coarse, and stiff in some cases, and greatly elongated and highly elastic in those species in which it has attained its fullest development . Its function is to See also:strain the water from the small marine molluscs, crustaceans, or fish upon which the whales subsist .

In feeding, whales fill the immense mouth with water containing shoals of these small creatures, and then, on closing the jaws and raising the See also:

tongue, so as to diminish the cavity of the mouth, the water streams out through the narrow intervals between the hairy fringe of the whalebone blades, and escapes through the lips, leaving the living prey to be swallowed . Although sometimes divided into two families, Balaenidae and Balaenopteridae, whalebone-whales are best included in a single See also:family group under the former name . The typical members of this family are the so-called right-whales, forming the genus Balaena, in which there are no folds on the See also:throat and See also:chest, and no back-fin; while the cervical vertebrae are fused into a single mass . The flippers are short and broad, with five digits; the head is very large and the whalebone very long and narrow, highly elastic and black; while the scapula is high, with a distinct coracoid and coronoid process . This genus contains the well-known Greenland right-whale (B. mysticetus) of the See also:Arctic seas, the whalebone and oil of which are so much valued in See also:commerce, and also other whales, distinguished by having the head somewhat smaller in proportion to the body, with shorter whalebone and a larger number of vertebrae . These inhabit the temperate seas of both See also:northern and See also:southern hemispheres, and have been divided into species in accordance with their See also:geographical See also:distribution, such as B. biscayensis of the See also:North See also:Atlantic, B. japonica of the North Pacific, B. australis of the South Atlantic, and B. antipodarum and novae-zelandiae of the South Pacific; but the See also:differences between them are so small that they may probably be regarded as races of a single species, the black whale (B. australis) . On the head these whales carry a peculiar structure which is known to whalers as the " See also:bonnet." This is a large horny excrescence, worn into hollows like a much-denuded piece of See also:limestone rock, growing probably in the neighbourhood of the blow-hole . More than one theory has been suggested to account for its presence . One See also:suggestion is that it indicates the descent of whales from See also:rhinoceros-like mammals; another that this species of whale is in the See also:habit of rubbing against rocks in order to free itself from barnacles, and thus produces a kind of See also:corn—although why on the See also:nose alone is not stated . Dr W . G . Ridewood, however, considers that the structure is due to the fact that the horny layers which are produced all over the skin are not See also:shed on this particular spot .

The pigmy whale (1eobalaena marginata) represents a genus agreeing with the right-whales in the See also:

absence of throat-flutings, and with the rorquals in the presence of a dorsal fin . The cervical vertebrae are united, and there are only 43 vertebrae altogether . The flippers are small, narrow, and with only four digits . The ribs remarkably expanded and flattened; the scapula low and broad, with completely developed acromion and coracoid processes . The whalebone is long, slender, elastic and white . The species which inhabits the South See also:American, Australian and New See also:Zealand seas is the smallest of the whalebone-whales, being not more than 20 ft. in length . In contrast to the preceding is the great See also:grey whale (Rachianectes See also:glaucus) of the North Pacific, which combines the relatively small head, elongated shape, and narrow flippers of the fin-whales, with the smooth throat and absence of a back-fin distinctive of the right-whales . The whalebone is shorter and coarser than in any otherspecies . In the See also:skeleton the cervical vertebrae are free, and the first two ribs on each side expanded and united to form a large bony See also:shield . In the humpback-whale (Megaptera longimana or boops) the head is of moderate size, the whalebone-plates are short and wide, and the cervical vertebrae free . The skin of the throat is fluted so as to form an expansible pouch; there is a low back-fin; and the flippers, which have four digits each, are extremely long, equalling about one-See also:fourth the See also:total length of the animal . The acromion and coracoid processes of the scapula are rudimentary .

See HUMPBACK-WHALE . The right-whales are built for cruising slowly about in See also:

search of the shoals of small floating invertebrates which form their food, and are consequently broad in See also:beam, with a See also:float-shaped body and immovable neck . The humpback is of somewhat similar build, but with a smaller head, and probably attains considerable See also:speed owing to the length of its flippers . The finners, or rorquals (Balaenoptera), which prey largely on fish, are built entirely for speed, and are the ocean greyhounds of the group . Their bodies are consequently long and attenuated, and their necks are partially See also:mobile; while they are furnished with capacious pouches for storing their food . They chiefly differ from the humpback by the smaller head, long and slender build, small, narrow, and pointed flippers, each containing four digits, and the large acromion and coracoid processes to the low and broad scapula . Rorquals are found in almost every See also:sea . Among them are the most gigantic of all animals, B. sibbaldi, which attains the length of 8o ft., and the small B. rostrata, which does not exceed 30 . There are certainly four distinct modifications of this genus, represented by the two just mentioned, and by B. musculus and B. borealis, all inhabitants of See also:British seas, but the question whether almost identical forms found in the See also:Indian, Southern and Pacific Oceans are to be regarded as specifically identical or as distinct awaits future researches, although some of these have already received distinct names . See RORQUAL . In the See also:report on the See also:zoology of the " See also:Discovery " expedition, published in 1907 by the British Museum, E . A .

See also:

Wilson describes a whale frequenting the fringe of the See also:Antarctic See also:ice which indicates a new generic type . Mainly black in colour, these whales measure about 20 or 30 ft. in length, and have a tall dorsal fin like that of a killer . Toothed Whales.—The second suborder is represented by the toothed whales, or Odontoceti, in which there is no whalebone, and teeth, generally numerous, though sometimes reduced to a single pair, and occasionally wanting, are normally developed . Unlike that of the whalebone-whales, the upper surface of the skull is more or less unsymmetrical . The nasal bones are in the form of nodules or flattened plates, applied closely to the frontals, and not forming any part of the roof to the nasal passage, which is directed upwards and backwards . The olfactory organ is rudimentary or absent . Hinder end of the maxilla expanded and covering the greater part of the orbital plate of the frontal bone . Lacrymal bone either in-separable from the jugal, or, if distinct, large, and forming part of the roof of the See also:orbit . Tympanic bone not welded with the periotic, which is usually only attached to the See also:rest of the skull by See also:ligament . Two halves of the lower jaw nearly straight, expanded in height posteriorly, with a wide See also:funnel-shaped aperture to the dental canal, and coming in contact in front by a See also:flat surface of variable length, but constituting a symphysis . Several of the anterior ribs with well-developed capitular processes, which articulate with the bodies of the vertebrae . Sternum almost always composed of several pieces, placed one behind the other, with which several pairs of ribs are connected by well-developed cartilaginous or ossified sternal ribs .

External respiratory aperture single, the two nostrils uniting before they reach the surface, usually in the form of a transverse sub-crescentic valvular aperture, situated on the top of the head . Flippers with five digits, though the first and fifth are usually little developed . No caecum, except in Platanista . The first family, Physeteridae, is typified by the sperm-whale, and characterized by the absence of functional teeth in the upper jaw; the lower teeth being various, and often much reduced in number . Bones of the skull raised so as to form an elevated prominence or See also:

crest behind the nostrils . Pterygoid bones thick, produced backwards, meeting in the middle line, and not involuted to form the outer See also:wall of the post-palatine air-sinuses, but simply hollowed on their outer side . Transverse processes of the arches of the dorsal vertebrae, to which the tubercles of the ribs are attached, ceasing abruptly near the end of the series, and replaced by processes on the body at a lower level, and serially homologous anteriorly with the heads of the ribs, and posteriorly with the transverse processes of the lumbar vertebrae . Costal cartilages not ossified . The first group, or Physeterinae, includes the sperm-whale itself, and is characterized by the presence of a full series of lower teeth, which are set in a groove in place of sockets, the groove being imperfectly divided by partial septa, and the teeth held in place by the strong, fibrous gum . No distinct lacrymal bone . Skull strikingly asymmetrical in the region of the nasal apertures, in consequence of the See also:left opening greatly exceeding the right in size . In the sperm-whale ((Physeter macrocephalus) the upper teeth are apparently of uncertain number, rudimentary and functionless, being embedded in the gum .

Lower jaw with from 20 ' to 25 teeth on each side, stout, conical, recurved and pointed at the apex 772 until they are worn, without enamel . Upper surface of the skull See also:

concave ; its posterior and lateral edges raised into a very high and greatly compressed semicircular crest or wall (fig . 2) . Zygomatic processes of jugal bones thick and massive . Muzzle greatly elongated, broad at the base, and gradually tapering to the apex . Lower taw exceedingly long and narrow, the symphysis being more than See also:half the length . Vertebrae: C 7, D 11, L 8, Ca 24; total 50 . See also:Atlas, or first vertebra, free; all the other cervical vertebrae united by their bodies and spines into a single mass . See also:Eleventh pair of ribs rudimentary . Head about one-third the length of the body; very massive, high and truncated, and rather compressed in front; owing its huge size and form mainly to the See also:accumulation of a mass of fatty tissue filling the large hollow on the upper surface of the skull and overlying the long muzzle . The single blow-hole is longitudinal, slightly S-shaped, and placed at the upper and anterior extremity of the head to the left side of the middle line . The opening of the mouth is on the under side of the head, considerably behind the end of the snout .

Flippers short, broad and truncated . Dorsal fin represented by a low protuberance . See SPERM-WHALE . In the lesser or pigmy sperm-whale (Cogia breviceps) there may be a pair of rudimentary teeth in the upper jaw, while on each side of the lower jaw there are from 9 to 12 rather long, slender, pointed and curved teeth, with a coat-in of enamel . Upper surface of the skull concave, with thick, raised, posterior and lateral margins, massive and rounded at their anterior terminations above the orbits . Muzzle not longer than the See also:

cranial position of the skull, broad at the base, and rapidly tapering to the apex . Zygomatic process of the jugal rod-like . Lower jaw with symphysis less than half its length . Vertebrae: C 7, D 13 or 14, L and Ca 30; total 50 or 51 . All the cervical vertebrae united by their bodies and arches . The head is about one-See also:sixth of the length of the body, and obtusely pointed in front; the mouth small and placed far below the apex of the snout; the blow-hole crescentic, and placed obliquely on the See also:crown of the head in advance of the eyes and to the left of the middle line; while the flippers are bluntly sickle-shaped, and the back-fin triangular . This species attains a length of from 9 to 13 ft .

Phoenix-squares

A second subfamily is represented by the See also:

bottle-noses and beaked whales, and known as the Ziphiinae . In this group the lower teeth are rudimentary and concealed in the gum, except one, or rarely two, pairs which may be largely developed, especially in the male . There is a distinct lacrymal bone . Externally the mouth is produced into a slender rostrum or beak, from above which the rounded See also:eminence formed by a See also:cushion of fat resting on the cranium in front of the blow-hole rises somewhat abruptly . The blow-hole is single, crescentic and median, as in the Delphinidae . Flippers small, ovate, with five digits moderately well developed . A small obtuse dorsal fin situated considerably behind the middle of the back . Longitudinal grooves on each side of the skin of the throat, diverging posteriorly, and nearly meeting in front . In external characters and habits the whal