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DISTRIBUTION OF

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Originally appearing in Volume V21, Page 783 of the 1911 Encyclopedia Britannica.
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DISTRIBUTION OF  See also:PLANTS See also:Common experience shows that temperature is the most important See also:condition which controls the See also:distribution of plants . Those of warmer countries cannot be cultivated in See also:British gardens without See also:protection from the rigours of See also:winter; still less are they able to hold their own unaided in an unfavourable See also:climate . Temperature, then, is the fundamental limit which nature opposes to the indefinite See also:extension of any one See also:species . See also:Buffon remarked " that the same temperature might have been expected, all other circumstances being equal, to produce the same beings in different parts of the globe, both in the See also:animal and See also:vegetable kingdoms." Yet lawns in the See also:United States are destitute of the common See also:English See also:daisy, the See also:wild See also:hyacinth of the See also:woods of the United See also:Kingdom is absent from See also:Germany, and the See also:foxglove from See also:Switzerland . We owe to Buffon the recognition of the See also:limitation of See also:groups of species to regions separated from one another by " natural barriers." When by the aid of See also:man they surmount these, they often dominate with unexpected vigour the native vegetation amongst which they are colonists . The See also:cardoon and See also:milk See also:thistle, both See also:European plants, See also:cover tracts of See also:country in See also:South See also:America with impenetrable thickets in which both man and beast may be hopelessly lost . The watercress blocks the See also:rivers of New See also:Zealand into which it has been introduced from See also:Europe . The problem, then, which plant-distribution presents is twofold: it has first to See also:map out the See also:earth's See also:surface into " regions " or " areas of vegetation," and secondly to trace the causes which have brought them about and led to their restriction and to their mutual relations . The earliest attempts to See also:deal with the first See also:branch of the inquiry may be called physiognomical . They endeavoured to define " aspects of vegetation " in which the " forms " exhibited an obvious See also:adaptation to their See also:climatic surroundings . This has been done with success and in See also:great detail by Grisebach, whose Vegetation der Erde from this point of view is still unsurpassed . With it may be studied with See also:advantage the unique collection at See also:Kew of pictures of plant-See also:life in its broadest aspects, brought together by the See also:industry and munificence of See also:Miss Marianne See also:North .

Grisebach declined to see anything in such " forms " but the See also:

production by nature of that which responds to See also:external conditions and can only exist as See also:long as they remain unchanged . We may agree with Schimper that such a point of view is obsolete without rejecting as valueless the admirable See also:accumulation of data of which it admittedly fails to give any rational explanation . A single example will be sufficient to illustrate this . The genus Senecio, with some loco species, is practically See also:cosmopolitan . In external See also:habit these exhibitadaptations to every See also:kind of climatic or See also:physical condition: they may be See also:mere weeds like groundsels or ragworts, or climbers masquerading like See also:ivy, or succulent and almost leafless, or they may be shrubs and even trees . Yet thrcughout they agree in the essential structure of their floral See also:organs . The cause of such agreement is, according to Grisebach, shrouded in the deepest obscurity, but it finds its obvious and See also:complete explanation in the descent from a common ancestor which he would unhesitatingly reject . From this point of view it is not sufficient, in attempting to map out the earth's surface into " regions of vegetation, " to have regard alone to adaptations to physical conditions . We are compelled to take into See also:account the actual See also:affinity of the plants inhabiting them . Anything See also:short of this is merely descriptive and empirical, and affords no rational basis for inquiry into the mode in which the distribution of plant-life has been brought about . Our regions will not be " natural " unless they See also:mark out real discontinuities both of origin and affinity, and these we can only seek to explain by reference to past changes in the earth's See also:history . We arrive thus at " the essential aim of See also:geographical See also:botany," which, as stated by Schimper, is " an inquiry into the causes of See also:differences existing among the various floras." To quote further: " Existing floras exhibit only one moment in the history of the earth's vegetation .

A transformation which is sometimes rapid, some-times slow, but always continuous, is wrought by the reciprocal See also:

action of the innate variability of plants and of the variability of the external factors . This See also:change is due partly to the migrations of plants, but chiefly to a transformation of the plants covering the earth." This transformation is due to new characters arising through variation . " If the new characters be useful, they are selected and perfected in the descendants, and constitute the so-called `adaptations' in which the external factors acting on the plants are reflected." The study of the nature of these adaptations, which are often extremely subtle and by no means merely superficial, is termed Ecology (see above) . The remark may conveniently find its See also:place here that plants which have reached a high degree of adaptive specialization have come to the end of their tether: a too complicated See also:adjustment has deprived them of the See also:elasticity which would enable them to adapt themselves to any further change in their surroundings, and they would pass away with conditions with which they are too inextricably See also:bound up . Vast floras have doubtless thus found their See also:grave in geologic change . That wrought by man in destroying forests and cultivating the See also:land will be no less effective, and already specimens in our herbaria alone represent species no longer to be found in a living See also:state . Extinction may come about indirectly and even more surely . This is easy to happen with plants dependent on See also:insects for their fertilization . Kronfeld has shown that aconites are dependent for this on the visits of a Bombus and cannot exist outside the See also:area where it occurs . The actual and past distribution of plants must obviously be controlled by the facts of physical See also:geography . It is concerned with the land-surface, and this is more symmetrically disposed than would at first sight appear from a glance at a map of the See also:world . See also:Lyell points out that the See also:eye of an observer placed above a point between See also:Pembroke and See also:Wexford, See also:lat .

52° N. and long . 6° W., would behold at one view the greatest possible quantity of land, while the opposite hemisphere would contain the greatest quantity of See also:

water . The See also:continental area is on one See also:side of the See also:sphere and the oceanic on the other . Love has shown (Nature, Aug . 1, 1907, p . 328) that this is the result of physical causes and that the existence of the Pacific Ocean " shows that the centre of gravity of the earth does not coincide with the centre of figure." One See also:half of the earth has therefore a greater See also:density than the other . But " under the See also:influence of the rotation the parts of greater density tend to recede further from the See also:axis than the parts of less density . . . the effect must be to produce a sort of furrowed surface." The furrows are the great ocean basins, and these would still persist even if the land surface were enlarged to the 1400 fathoms See also:contour . These considerations preclude the possibility of solving difficulties in geographical distribution by the construction of hypothetical land-surfaces, an expedient which See also:Darwin always stoutly opposed (Life and Letters, ii . 74-78) . The furrowed surface of the earth gives the land-area a See also:star-shaped figure, which may from See also:time to time have varied in outline, but in the See also:main has been permanent . It is excentric as regards the See also:pole and sends tapering extensions towards the south .

See also:

Sir See also:George Darwin finds a possible explanation of these in the screwing See also:motion which the earth would suffer in its plastic state . The polar regions travelled a little from See also:west to See also:east relatively to the See also:equatorial, which lagged behind . The great See also:primary divisions of the earth's See also:flora See also:present them-selves at a glance . The tropics of See also:Cancer and Capricorn cut off with surprising precision (the latter somewhat less so) the tropical from the north and south temperate zones . The north temperate region is more sharply separated from the other two than the south temperate region from the tropical . I . NORTH TEMPERATE REGION (Holarctic).—This is the largest of all, circumpolar, and but for the break at See also:Bering Straits, would be, as it has been in the past, continuous in both the old and new worlds . It is characterized by its See also:needle-leaved Coniferae, its catkin-bearing (Amentaceae) and other trees, See also:deciduous in winter, and its profusion of herbaceous species . II . SOUTH TEMPERATE REGION.—This occupies widely separated areas in South See also:Africa, See also:Australia, New Zealand and South America . These are connected by the presence of See also:peculiar types, Proteaceae, Restiaceae, Rutaceae, &c., mostly shrubby in habit and on the whole somewhat intolerant of a moist climate . Individual species are extremely numerous and often very restricted in area .

A See also:

consideration of these regions makes it apparent that they are to a large extent adaptive . The boreal is See also:cold, the austral warm, and the tropical affords conditions of See also:heat and moisture to which the vegetation of the others would be intolerant . If we take with Drude the number of known families of flowering plants at 240, 92 are generally dispersed, 17 are more restricted, while the See also:remainder are either dominant in or peculiar to See also:separate regions . Of these 40 are boreal, 22 austral and 69 tropical . If we add to the latter figure the families which are widely dispersed, we find that the tropics possess 161 or almost exactly two-thirds of the large groups comprised in the world's vegetation . M . Casimir de See also:Candolle has made an See also:independent investigation, based on See also:Hooker and See also:Bentham's Genera pientarum . The result is unfortunately (1910) unpublished, but he informs the present writer that the result leads to the striking conclusion: " La vegetation est un phenomene surtout intertropical, dont nous ne voyons plus que restes affaiblis dans nos regions teinperees." In attempting to account for the distribution of existing vegetation we must take into account palaeontological See also:evidence . The results arrived at may be read as a sequel to the See also:article on See also:PALAEOBOTANY . The vegetation of the Palaeozoic era, till towards its See also:close, was apparently remarkably homogeneous all over the world . It was characterized by arborescent vascular Cryptogams and See also:Gymnosperms of a type (Cordaiteae) which have See also:left no descendants beyond it . In the See also:southern hemisphere the Palaeozoic flora appears ultimately to have been profoundly modified by a lowering of temperature and the existence of glacial conditions over a wide area .

It was replaced by the Glossopteris flora which is assumed to have originated in a vast continental area (See also:

Gondwana land), of which remnants remain in South America, South Africa and Australia . The Glossopteris flora gradually spread to the See also:northern hemisphere and intermingled with the later Palaeozoic flora which still persisted . Both were in turn replaced by the See also:Lower Mesozoic flora, which again is thought to have had its See also:birth in the hypothetical Gondwana land, and in which Gymnosperms played the leading See also:part. formerly taken by vascular Cryptogams . The abundance of Cycadean plants is one of its most striking features . They attained the highest degree of structural complexity in the Bennettiteae, which have been thought even to foreshadow a floral organization . Though now on the way to extinction, Cycadeae are still widely represented in the southern hemisphere by genera which, however, have no counterpart in the Mesozoic era . Amongst Conifers the archaic genera, Ginkgo and See also:Araucaria still persist . Once widely distributed in the See also:Jurassic See also:period throughout the world, they are now dying out: the former is represented by the solitary See also:maiden-See also:hair See also:tree of See also:China and See also:Japan; the latter by some ten species confined to the southern hemisphere, once perhaps their See also:original See also:home . With them may be associated the anomalous Sciadopitys of Japan . So far the See also:evolution of the vegetable kingdom has proceeded with-out any conspicuous break . Successive types have arisen in ascending sequence, taken the See also:lead, and in turn given way to others . Butthe later period of the Mesozoic era saw the almost sudden See also:advent of a fully See also:developed angiospermous vegetation which rapidly occupied the earth's surface, and which it is not easy to See also:link on with any that preceded it .

The closed ovary implies a mode of fertilization which is profoundly different, and which was probably correlated with a simultaneous development of See also:

insect life . This was accompanied by a vegetative organization of which there is no obvious foreshadowing . As See also:Clement See also:Reid remarked: " World-wide floras, such as seem to characterize some of the older periods, have ceased to be, and plants are distributed more markedly according to geographical provinces and in climatic zones." The See also:field of evolution has now been transferred to the northern hemisphere . Though See also:Angiosperms become dominant in all known plant-bearing Upper Cretaceous deposits, their origin See also:dates even earlier . In Europe Heer's Populus primaeva from the Lower Cretaceous in See also:Greenland was long accepted as the See also:oldest dicotyledonous plant . Other undoubted See also:Dicotyledons, though of uncertain affinity, of similar See also:age have now been detected in North America . The Cenomanian rocks of Bohemia have yielded remains of a sub-tropical flora which has been compared with that existing at present in Australia . Upper Cretaceous formations in America have yielded a copious flora of a warm-temperate climate from which it is evident that at least the generic types of numerous not closely related existing dicotyledonous trees had already come into existence . It may be admitted that the See also:identification of fragmentary See also:leaf-remains is at most See also:precarious . Even, however, with this See also:reservation, it is difficult to resist the See also:mass of evidence as a whole . And it is a plausible conjecture that the vegetation of the globe had already in its main features been constituted at this period much as it exists at the present moment . There were oaks, beeches (scarcely distinguishable from existing species), birches, planes and willows (one closely related to'the living Salix candida), laurels, represented by Sassafras and Cinnamomum, magnolias and See also:tulip trees (Liriodendron), myrtles, See also:Liquidambar, Diospyros and ivy .

This is a flora which, thinned out by losses, practically exists to this See also:

day in the southern United States . And one essentially similar but adapted to slightly cooler conditions existed as far north as the See also:latitude of Greenland . The See also:tertiary era opens with a climate in which during the See also:Eocene period something like existing tropical conditions must have obtained in the northern hemisphere . The remains of palms (Sabal and Nipa) as well as of other large-leaved Monocotyledons are preserved . See also:Sequoia (which had already appeared in the See also:American Upper Cretaceous) and the deciduous See also:cypress (Taxodium distichum) are found in Europe . Starkie See also:Gardner has argued with much plausibility that the Tertiary floras which have been found in the far north must have been of Eocene age . That of See also:Grinnell Land in lat . 81 ° consisted of Conifers (including the living spruce), poplars and willows, such as would be found now 25° to the south . The flora of Disco See also:Island in lat . 700 contained Sequoia, planes, maples and magnolias, and closely agrees with the See also:Miocene flora of central Europe . Of this copious remains have been found in Switzerland and have been investigated with great ability by O . Heer .

They point to cooler conditions in the northern hemisphere: palms and tropical types diminish; deciduous trees increase . Sequoia and the tulip-tree still remain; See also:

figs are abundant; laurels are represented by Sassafras and camphor; herbaceous plants (See also:Ranunculaceae, See also:Cruciferae, See also:Umbelliferae) are present, though, as might be expected, only fragmentarily preserved . We may draw with some certainty the conclusion that a See also:general See also:movement southward of vegetation had been brought about . While Europe and probably North America were occupied by a warm temperate flora, tropical types had been driven southward, while the adaptation of others to See also:arctic conditions had become accentuated . A See also:gradual refrigeration proceeded through the See also:Pliocene period . This was accompanied in Europe by a drastic weeding out of Miocene types, ultimately leaving the flora See also:pretty much as it now exists . This, as will be explained, did not take place to anything like the same extent in North America, the vegetation of which still pre-serves a more Miocene facies . Torreya, now confined to North America and Japan, still lingered,- as did Ocotea, now profusely developed in the tropics, but in north temperate regions only existing in the Canaries: the See also:evergreen oaks, so characteristic of the Miocene, were reduced to the existing Quercus ilex . At the close of the Pliocene the European flora was apparently little different from that now existing, though some warmer types such as the water-See also:chestnut (Trapa natans) had a more northern extension . The glacial period effected in Europe a wholesale extermination of temperate types accompanied by a southern extension of the arctic flora . But Its operation was in great measure See also:local . The Pliocene flora found refuges in favoured localities from which at its close the lowlands were restocked while the arctic plants were left behind on the mountains .

During the milder interglacial period some southern types, such as See also:

Rhododendron ponticum, still held their own, but ultimately succumbed . The evidence which has thus been briefly summarized, points unmistakably to the conclusion that existing vegetation originated in the northern hemisphere and under climatic conditions corresponding to what would now be termed sub-tropical . It occupied a continuous circumpolar area which allowed See also:free communication between the old and new worlds . The gradual differentiation of their floras has been brought about rather by extermination than specialization, and their distinctive facies by the development and multiplication of the surviving types . The distribution of See also:mountain barriers in the Old and New Worlds is in striking contrast . In the former they run from east to west; in the latter from north to south . In the Old World the boreal See also:zone is almost sharply cut off and afforded no means of See also:escape for the Miocene vegetation when the climate became more severe . Thus in the Mediterranean region the large groups of palms, figs, myrtles and laurels are each only represented by single surviving species . The great tropical See also:family of the Gesneraceae has left behind a few outliers: Ramondia in the See also:Pyrenees, Haberlea in the Balkans, and Jankaea in See also:Thessaly; the Pyrenees also possess a See also:minute Dioscorea, See also:sole European survivor of the yams of the tropics . In North America there is no such barrier: the Miocene flora has been able to escape by See also:migration the fluctuations of climate and to return when they ameliorated . It has preserved its characteristic types, such as See also:Magnolia, Liriodendron, Liquidambar, Torreya, Taxodium and Sequoia . While it has been customary to describe the Miocene flora of Europe as of a North American type, it would be more accurate to describe the latter as having in great measure preserved its Miocene See also:character .

If mountains serve as barriers which See also:

arrest the migration of the vegetation at their See also:base, their upper levels and summits afford lines of communication by which the floras of colder regions in the northern hemisphere can obtain a southern extension even across the tropics . They doubtless equally See also:supply a path by which southern temperate types may have extended northwards . Thus the characteristic assemblage of plants to which Sir See also:Joseph Hooker has given the name Scandinavian " is present in every latitude of the globe, and is the only one that is so " (Trans . Linn . See also:Soc. See also:xxiii . 253) . In the mountains of See also:Peru we find such characteristic northern genera as Draba, Alchemilla, Saxifraga, Valeriana, Gentiana and Bartsia . High elevations reproduce the physical conditions of high latitudes . The aqueous vapour in the See also:atmosphere is transparent to luminous but opaque to obscure heat-rays . The latter are retained to warm the See also:air at lower levels, while it remains cold at higher . It results that besides a See also:horizontal distribution of plants, there is also an altitudinal: a fact of See also:cardinal importance, the first observation of which has been attributed to See also:Tournefort . Speaking generally, all plants tend to exhaust particular constituents of the See also:soil on which they grow .

Nature therefore has provided various contrivances by which their seeds are disseminated beyond the actual position they occupy . In a large number of cases these only provide for migration within sufficient but narrow limits; such plants would be content to remain local . But other physical agencies come into See also:

play which may be briefly noticed . The first of these is See also:wind: it cannot be doubted that small seeds can be swept I herbacea, Silene acaulis and Dryas octopetala will serve as examples. up like dust and transported to considerable distances . This is Even so small an area as that of See also:Britain illustrates what has already certainly the See also:case with See also:fern-spores . The vegetation of See also:Krakatoa been pointed out, that the species of a flora change both with latitude and See also:altitude . See also:Watson further brought out the striking fact that the west and east of Britain each had species peculiar to it; the former he characterized as See also:Atlantic, the latter as Germanic . The Cornish See also:heath (Erica vagans) and the maiden-hair fern (Adiantum Capillus-Veneris) may serve as instances of the one, the man-orchis (Aceras anthropophora) and Reseda lutea of the other . See also:Ireland illustrates the same fact . It possesses about moo species, or about two-thirds the number of Britain . On its western shores there are some twenty, such as Saxifraga umbrosa, Erica mediterranea and Arbutus unedo, which are not found in Britain at all . The British Phanerogamic flora, it may be remarked, does not contain a single endemic species, and 38 % of the See also:total number are common to the three northern continents .

The See also:

analysis of larger areas yields results of the same kind . Within the same region we may expect to find considerable differences as we pass from one See also:meridian to another . Assuming that in its circumpolar origin the North Temperate flora was fairly homogeneous, it would meet in its centrifugal extension with a wide range of local conditions; these would favour the preservation of numerous species in some genera, their greater or less elimination in others . Thus comparing the Nearctic and Palaearctic floras we find striking differences overlying the points of agreement already indicated . The former is poor in Cruciferae, See also:Caryophyllaceae, Umbelliferae, See also:Primulaceae and See also:Labiatae; but for the occurrence of Calluna in Newfound-land it would have no heaths . On the other See also:hand, it is See also:rich in See also:Compositae, especially Solidago and See also:Aster, Polemoniaceae, Asclepiadaceae, Hydrophyllaceae and See also:Cyperaceae, and it has the endemic See also:Sarracenia, type of a family structurally allied to poppies, of which of the remaining genera See also:Darlingtonia is Californian, and Heliamphora Venezuelan . These distinctions led Sir Joseph Hooker to claim for the two divisions the See also:rank of primary regions . Darwin doubted, however, whether they ought to be separated (Life, iii . 230) . Lyell, discussing the facts of zoological distribution, admits that " the farther we go north . . . the more the discordance in genera and species diminishes " (Principles, ii . 340); and Hemsley finds that not less than 75 % of the genera in the flora of eastern North America " are represented in the old world," and, according to See also:Asa See also:Gray, 50 % in Europe .

Latitudinally the region subdivides naturally into several well-marked sub-regions which must be briefly discussed . as in the Old World—oaks, chestnuts, beeches, hazels, hornbeams, birches, alders, willows and poplars . Or to take the small but well-defined See also:

group of five-leaved pines, all the species of which may be seen growing side by side at Kew under identical conditions: we have the See also:Weymouth See also:pine (Pinus Strobus) in eastern North America, P. monticola and the See also:sugar pine (P . Lambertiana) in See also:California, P . Ayacahuste in See also:Mexico, the Arolla pine (P . Cembra) in Switzerland and See also:Siberia, P . Pence in See also:Greece, the Bhotan pine (P. excelsa) in the Himalayas, and two other species in Japan . Amongst broad-leaved trees Juglans has a similar Holarctic range, descending to the West Indies; so has Aesculus, were it not lacking in Europe; it becomes tropical in South America and Malaya . If we turn to herbaceous plants, Hemsley has pointed out that of the thirteen genera of Ranunculaceae in California, eleven are British . While the tropics preserve for us what remains of the pre-Tertiary or, at the latest, Eocene vegetation of the earth, which formerly had a much wider extension, the flora of the North Temperate region is often described as the survival of the Miocene . Engler therefore calls it Arcto-Tertiary . We must, however, agree with Starkie Gardner that it is only Miocene as regards its present position, which was originally farther north, and that its actual origin was much earlier .

There has been in effect a successive shifting of zones of vegetation southwards from the pole . Their distinctive and adaptive characteristics doubtless began to be established as soon as the phanerogamic flora was constituted . There is no See also:

reason to suppose that the peculiarities of the arctic flora are more See also:modern than those of any other, though there is no fossil evidence to prove that it was not so . The North Temperate region admits of subdivision into several well-marked sub-regions . The general method by which this is effected in this and other cases is statistical . As A. de Candolle, however, points out, exclusive reliance on this may be misleading unless we also take into account the character and See also:affinities of the plants dealt with (Geogr . Bot. i . 1166) . The numerical predominance of certain families or their See also:absence affords criteria for marking out boundary lines and tracing relationships . The analysis of the flora of the British Isles will afford an See also:illustration . This was first attempted in 1835 by H . C .

Watson, and his conclusions were en-forced ten years later by See also:

Edward See also:Forbes, who dealt also with the See also:fauna . Watson showed that See also:Scotland primarily, and to a less extent the north of See also:England, possessed species which do not reach the south . Such are the See also:crowberry (Empetrum nigrum), Trientalis europaea, Rubus saxatilis and the globe-See also:flower (Trollius europaeus) . He further found that there was an See also:element which he termed " boreal in a more intense degree," which amounted to about " a fifteenth of the whole flora." This was not confined to the north .but may occur on the mountains of England and See also:Wales: Salix was completely exterminated in 1883 by a thick coat of red-hot See also:pumice . Yet in 1886 Treub found that it was beginning to cover itself again with plants, including eleven species of ferns; but the nearest source of supply was to m. distant . Seeds are carried with more facility when provided with plumes or wings . Treub found on Krakatoa four species of composites and two See also:grasses . Water is another obvious means of transport . The littoral vegetation of See also:coral islands is derived from See also:sea-See also:borne fruits . The seeds of West See also:Indian plants are thrown on the western shores of the British Isles, and as they are capable of germination, the species are only pre-vented from establishing themselves by an uncongenial climate . Travers picked up a See also:seed of Edwardsia in the See also:Chatham Islands, evidently washed ashore from New Zealand (Linn . Soc .

Journ. ix . 1865) . Rivers bring down the plants of the upper levels of their basins to the lower: thus species characteristic of the See also:

chalk are found on the See also:banks of the See also:Thames near See also:London . Birds are even more effective than wind .in transporting seeds to long distances . Seeds are carried in soil adhering to their feet and plumage, and aquatic plants have in consequence for the most part an exception-ally wide range . See also:Fruit-pigeons are an effective means of transport in the tropics by the undigested seeds which they void in their excrement . Quadrupeds also play their part by carrying seeds or fruits entangled in their coats . Xanthium spinosum has spread from the See also:Russian See also:steppes to every stock-raising country in the world, and in some cases has made the industry impossible . Even insects, as in the case of South See also:African locusts, have been found to be a means of distributing seeds . The primary regions of vegetation, already indicated, and their subordinate provinces may now be considered more in detail . I . NORTH TEMPERATE REGION.—Many writers on the distribution of animals prefer to separate this into two regions of " primary rank " : the Palaearctic and the Nearctic .

But to justify such a See also:

division it is necessary to establish either an exclusive See also:possession or a marked predominance of types in the one which are correspondingly deficient in the other . This cannot apparently be done for insects or for birds; See also:Newton accordingly unites the two into the Holarctic region . It equally fails for plants . To take, for example, one of the most characteristic features of the Palaearctic region, its catkin-bearing deciduous trees: in North America we find precisely the same genera 1 . The Arctic-Alpine sub-region consists of races of plants belonging originally to the general flora, and recruited by subsequent additions, which have been specialized in See also:low stature and great capacity of endurance to survive long dormant periods, sometimes even unbroken in successive years by the transitory activity of the brief summer . It is continuous See also:round the pole and roughly is bounded by the arctic circle . Mature seeds are highly tolerant of cold and have been shown to be capable of withstanding the temperature even of liquid See also:hydrogen . Arctic plants make their brief growth and flower at a temperature little above freezing point, and are dependent for their heat on the See also:direct rays of the See also:sun . Characteristic representatives are Pa paver nudicaule, Sazifraga oppositifolia, which forms a profuse See also:carpet, and Dryas octopetala . Such plants perhaps extend to the most northern lands at present known . On May 3oth, in See also:Ward See also:Hunt's Island, lat . 83° 5', Sir George See also:Nares found that vegetation was fairly represented as regards quantity in the See also:poppy, See also:saxifrage and small tufts of grass." We may compare this with extreme alpine conditions: on a spot above the Aletsch See also:glacier at a height of 10,700 ft .

See also:

Ball found the temperature one See also:inch below the surface to be 83°, and he collected " over See also:forty species in flower." Taking the whole arctic flora at 762 species, Hooker found that 616 occurred in arctic Europe, and of these 586 are Scandinavian . Beyond the arctic circle some 200, or more than a See also: