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See also:EGG (O.E. aeg, cf. Ger. Ei, Swed. aegg, and prob. Gr. u,ov, See also:Lat. ovum)
, the See also:female reproductive See also:cell or ovum of animals, which gives rise generally only after fertilization to the See also:young
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The largest eggs are those of birds; and this because, to the minutes essential portion of the See also:egg, or germ, from which the young See also:bird grows, there is added a large See also:store of See also:food-material--the yolk and See also:
Though in shape differing but little from that of the See also:familiar See also:hen's egg, certain well-marked modifications of form are yet to be met with
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Thus the eggs of the See also:plover are See also:pear-shaped, of the See also:sand-See also:grouse more or less cylindrical, of the owls and titmice spherical and of the grebes biconical
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In the See also:matter of coloration the eggs of birds present a remark-able range
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The See also:pigments to which this coloration is due have been shown, by means of their absorption spectra (Sorby, Proc
.
Zool
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See also:Soc., 1875), to be seven in number
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The first of these, oorhodeine, is See also: That the egg during this passage turns slowly on its See also:long See also:axis is shown by the fact that the spots and lines have commonly a See also:spiral direction; though some of the markings are made during periods of See also:rest, as is shown by their See also:sharp outlines, See also:movement giving a blurred effect . Where the egg is pyriform, the large end makes way for the smaller . Many eggs display, in addition to the strongly marked spots, more or fewer fainter spots embedded in a deeper layer of the shell, and hence such eggs'are said to be " See also:double-spotted!" e.g. rails and plovers . Among some species, as in birds of See also:prey, the intensity of this coloration is said to increase with See also:age up to a certain point, when it as gradually decreases . Frequently, especially where but two eggs are laid (See also:Newton), all the dye will be deposited, sometimes on the first, sometimes on the last laid, leaving the other colour-less . But although of a number of eggs in a " clutch "—as the full See also:complement of eggs in a See also:nest is called—no two are exactly alike, they commonly See also:bear a very See also:close resemblance . Among certain species, however, which See also:lay several eggs, one of the number invariably differs markedly from the rest, as for example in the eggs of the See also:house-See also:sparrow or in those of the sparrow-See also:hawk, where, of a clutch of six, two generally differ conspicuously from the rest . Differing though these eggs do from the rest of the clutch, all yet present the characters See also:common to the species.' But the eggs of some birds, such as the Australian swamp See also:quail, Synoecus australis, present a remarkably wide range of variation in the matter of coloration, no two clutches being alike, the extremes ranging from pure white to eggs having a greenish ground colour and rufous spots or blotches . But a still more interesting See also:illustration of variation equally marked is furnished by the chikor See also:partridge (Caccabis chukar), since here the variation appears to be correlated with the See also:geographical See also:distribution of the species . Thus eggs taken in See also:Greece are for the most See also:part cream-coloured and unspotted; those from the Grecian See also:Archipelago are generally spotted and blotched; while more to the eastward spots are invariably present, and the blotches attain their maximum development . But in variability the eggs of the See also:guillemot (Lomvia troile) exceed all others: both in the See also:hue of the ground colour and in the form of the superimposed markings, these eggs exhibit a wonderful range for which no adequate explanation has yet been given . Individual peculiarities of coloration are commonly reproduced, not only with this species but also in others, See also:year after year . The coloration of the egg bears no sort of relation to the coloration of the bird which See also:lays it; but' it bears on the other signori. See also:hand a more or less See also:direct relation to the n.cure of the canceof environment during See also:incubation . colour . White eggs may generally be regarded as representing the See also:primitive type of egg, since they agree in this particular with the eggs of See also:reptiles . And it will generally be found that eggs of this hue are deposited in holes or in domed nests . So long indeed as nesting-places of this See also:kind are used will the eggs be white . And this because coloured eggs would be invisible in dimly lighted See also:chambers of this description, and therefore constantly exposed to the See also:risk of being broken by the sitting bird, or See also:rolling out of reach where the chamber was large enough to admit of this, whereas white eggs are visible so long as they can be reached by the faintest rays of See also:light . Pigeons invariably lay white eggs; and while some See also:deposit them in holes others build an open nest, a See also:mere See also:platform of sticks . These exceptions to the See also:rule show that the depredations of egg-eating animals are sufficiently guarded against by the overhanging foliage, as well as by the See also:great distance from the ground at which the nest is built . Birds which have reverted to the more See also:ancient See also:custom of nesting in holes after having See also:developed pigmented eggs, have adopted the See also:device of covering the shell with a layer of chalky matter (e.g. puffins), or, to put the case more correctly, they have been enabled to maintain survival after their return to the more ancient mode of See also:nidification, because this reversion was accompanied by the tendency to See also:cover the pigmented surface of the shell with this light-reflecting chalky incrustation . Eggs which are deposited on the See also:bare ground, or in other exposed situations, are usually protectively coloured: that is to say, the hue of the shell more or less completely harmonizes with the ground on which the egg is placed . The eggs of the plover tribe afford the most striking examples of this fact . But the majority of birds deposit their eggs in a more or less elaborately constructed nest, and `in such cases the egg, so far from being protectively coloured, often displays tints that would appear calculated rather to attract the See also:attention of egg-stealing animals; See also:bright blue or blue spotted with See also:black being commonly met with . It may be, however, that coloration of this kind is less conspicuous than is generally supposed, but in any case the safety of the egg depends not so much on its coloration as on the character of the nest, which, where protective devices are necessary, must harmonize sufficiently with its surroundings to See also:escape observation from prowling egg-stealers of all kinds . The See also:size of the egg depends partly on the number produced and partly on the conditions determining the See also:state of the young bird at hatching: hence there is a great disparity in the relative sizes of the eggs of different birds . Thus it will be found that young birds which emerge in the See also:world See also:blind, naked and helpless are the product of relatively small eggs, while on the contrary young hatched from relatively large eggs are down-clad and active from See also:birth . The fact that the eggs must be brooded. by the parent is also a controlling See also:factor in so far as number is concerned, for no more can be hatched than can be covered by the sitting bird . Other factors, however, less understood, also exercise a controlling See also:influence in this matter . Thus the ostrich lays from 12 to 16, the See also:teal 15, the partridge i2-20, while among many other species the number is strictly limited, as in the case of the hornbills and guillemots, which lay but a sLgle egg; the apteryx, See also:divers, petrels and pigeons never lay more than 2, while the gulls and plovers never exceed 4 . Tropical species are said to lay fewer eggs than their representatives in temperate regions, and further immature birds lay more and smaller eggs than when fully adult . Partly owing to the uniformity of shape, size and texture of the shell, the eggs of birds are by no means easy to distinguish, except in so far as their See also:family resemblances are concerned: that is to say, except in particular cases, they cannot be specifically distinguished, and hence they are of but little or no value for the purposes of See also:classification . See also:Save only among the megapodes, all birds brood their eggs, the See also:period of incubation varying from 13 days, as in small passerine birds, to 8 See also:weeks, as in the See also:cassowary, though eggs of the See also:rhea and of Struthio See also:hatch in from 5 to 6 weeks . But the megapodes deposit their eggs in mounds of decaying See also:vegetable matter or in sand in the neighbourhood of hot springs, and there without further apparent care leave them . Where the nestling is active from the moment of hatching the eggs have a relatively longer incubation period than in cases where the nestlings are for a long while helpless . Eggs of Mammals.—Only in the spiny See also:ant-eater, or See also:Echidna, and the See also:duck-billed See also:platypus, or Ornithorhynchus, among the See also:Mammalia, are the eggs provided with a large store of yolk, enclosed within a shell, and extruded to undergo development apart from the maternal tissues . In the case of the echidna the eggs, two in number, are about as large as those of a sparrow, similar in shape, and have a white, See also:parchment-like shell . After See also:expulsion they are transferred by the See also:beak of the See also:mother to a pouch resembling that of the marsupial kangaroos, and there they undergo development . The Otnithorhynchus, on the other hand, lays from two to four eggs, which in size and general See also:appearance resemble those of the echidna . They are, how-ever, deposited in a loosely constructed nest at the end of a long burrow and there brooded . In Marsupials, the eggs are smaller than those of Echidna and Ornithorhynchus, and they contain a larger proportion of yolk than occurs in higher mammals . Eggs of Reptiles.—The eggs of reptiles are invariably provided with a large amount of food yolk and enclosed with a See also:firm test or shell, which though generally parchment-like in texture may be calcareous as in birds, as, for example, in many of the tortoises and turtles and in the crocodiles . Among reptiles the egg is always white or yellowish, while the number laid often far exceeds that in the case of birds . The tuatara of New See also:Zealand, however, lays but ten—white hard-shelled, long and See also:oval—at intervals between See also:November and See also:January . The long intervals between the appearance of the successive eggs is a characteristic feature of the reptiles, but is met with among the birds only in the megapodes, which, like the reptiles, do not " brood " their eggs . Among the Chelonia the number of eggs varies from two to four in some of the tortoises, to aoo in some of the turtles: while in the crocodiles between 20 and 30 are produced, hard-shelled and white . The eggs of the lizards are always white or yellowish, and generally soft-shelled; but the geckos and the green See also:lizard lay 'hard-shelled eggs . Many of the soft-shelled eggs are remarkable for the fact that they increase in size after extrusion, owing to the stretching of the membranous shell by the growing embryo . In the matter of number lizards are less prolific than many of the Chelonia, a dozen eggs being the general number, though as many as See also:thirty may be produced at a See also:time, as in the case of the common See also:chameleon . While as a general rule the eggs of lizards are laid in burrows or buried, some are retained within the body of the parent until the young are ready to emerge; or they may even hatch within the oviduct . This occurs with some chameleons and some lizards, e. g. the slow-See also:worm . The common See also:English lizard is also viviparous . Normally the young leaves the egg immediately after its extrusion, but if by any See also:chance this extrusion is delayed they escape while yet in the oviduct . The majority of the See also:snakes lay eggs, but most of the vipers and the aquatic snakes are viviparous, as also are a few terrestrial species . The shell of the egg is always soft and parchment-like . As a rule the number of eggs produced among the snakes is not large, twenty or thirty being common, but some species of See also:python lay as many as a See also:hundred . Generally, among the oviparous snakes the eggs are buried, but some species of boas jealously guard them, enclosing them within the coils of the body . Eggs of See also:Amphibia.—Among the amphibia a greater variety obtains in the matter of the investment of the egg, as well as in the number, size and method of their disposal . The outer covering is formed by a toughening of the surface of a thick gelatinous coat which surrounds the essential parts of the egg . This coat in many species of See also:salamander—using this name in the wide sense—is produced into threads which serve either to See also:anchor the eggs singly or to bind them together in bunches . Viviparity occurs both among the limbless and the tailed Amphibia, the eggs hatching before they leave the oviduct or immediately after extrusion . The number of young so produced is generally not large, but the common salamander (Salamandra maculosa) may produce as many as fifty at a birth, though fifteen is the more normal figure . When the higher number is reached the young are relatively small and weak . As a rule among the Amphibia the young leave the egg in the form of larvae, generally known as " tadpoles "; but many I species produce eggs containing a sufficient amount of food material to enable the whole of the larval phase to be completed before hatching . Among the tailless Amphibia (frogs and toads) 'there are wide See also:differences in the number of eggs produced, while the methods by which these eggs are disposed of present a marvellous variety . As a rule vast quantities of eggs are See also:shed by the female into the See also:water in the form of " spawn." In the common See also:toad as many as 7000 eggs may be extruded at a time . These leave the body in the form of two long strings—one from each oviduct—of trans-lucent globules, gelatinous in texture, and enclosing a central See also:sphere of yolk, the upper See also:pole of which is black . The spawn of the common See also:frog differs from that of the toad in that the eggs all adhere to form a huge jelly-like See also:mass . But in many species the number of eggs produced are few; and these may be sufficiently stored with food-yolk to allow of the See also:tadpole See also:stage being passed before hatching, as in frogs of the genus Hylodes. in many cases the eggs are deposited out of the water and often in quite remarkable ways . Eggs of Fishes.—The eggs of fishes present an extremely wide range of form, and a no less extensive range in the matter of number . Both among the cartilaginous and bony fishes viviparity occurs . Most of the sharks and rays are viviparous, but in the oviparous species the eggs present some interesting and peculiar forms . Large in size, the outer coat or " shell " is in all cases See also:horn-like and flexible, but differs greatly in shape . Thus in the egg of the larger spotted See also:dog-See also:fish it is oblong in shape, flattened from See also:side to side, and has the angles produced into long, slender tendrils . As the egg is laid the lower tendrils project from the vent, and the mother rubs herself against some fixed body . The tendrils soon catch fast in some slight See also:projection, when the egg is dragged forth there to remain till hatching takes See also:place . A couple of narrow slits at each corner of the upper end serve to admit fresh water to the imprisoned embryo during the later stages of development; when development is See also:complete escape is made through the end of the shell . In the rays or " skates," long spines take the place of tendrils, the egg simply resting at the bottom of the See also:sea . The empty egg-cases of the rays are often found on the seashore, and are known as "Mermaids' purses." The egg of the See also:Port See also:Jackson See also:shark (Cestracion) is of enormous size, pear-shaped, and provided with a spiral flange extending along the whole length of the See also:capsule . In the See also:Chimaera the egg is long, more or less spindle-shaped, and produced on each side into a broad flange having a fringed edge, so that the whole bears a close resemblance to a long See also:leaf, broad and notched at one end, pointed at the other . This likeness to the seaweed among which it rests is doubtless a protective device, akin to that of protectively coloured birds' eggs . Among the bony fishes the eggs generally take the form of small See also:spheres, enclosed within a tough membrane or capsule . But they present many important differences, being in some fishes heavy and remaining at the bottom of the water, in other light and floating on the surface . While in some species they are distributed separately, in others they adhere together in masses . The eggs of the See also:salmon, for example, are heavy, hard and smooth, and deposited separately in a trough dug by the parent and afterwards covered to prevent them from being carried away by the stream . In the See also:perch they are adhesive and form long See also:band-like masses of spawn adhering to water-See also:plants . In the gobies the egg is spindle-shaped, and attached by one end by means of a network of See also:fibres, resembling rootlets; while in the See also:smelt the egg is loosely suspended by a membrane formed by the peeling off of a part of the outer sheath of the capsule . The eggs of the garfish (Belone vulgaris) and of the flying-fish of the genus Exocoetus, attach themselves to See also:foreign See also:objects, or to one another, by means of threads or cords developed at opposite poles of the egg . Among a number of fishes the eggs See also:float at the surface of the sea, often in enormous masses, when they are carried about at the See also:mercy of tides'and currents . An See also:idea of the size which such masses attain may be gathered from the fact that the spawn of the See also:angler-fish, Lophius piscatorius, takes the form of a See also:sheet from 2 to 3 ft. wide, and 30 ft. long . Another remarkable feature of these floating eggs is their transparency, inasmuch as they are extremely difficult to see, and hence they probably escape the rapacious maws of spawn-eating animals . The See also:cod tribe and fiat-fishes lay floating eggs of this description . The maximum number of eggs laid by fishes varies greatly, some species laying relatively few, others an enormous number . But in all cases the number increases with the See also:weight and age of the fish . Thus it has been calculated that the number laid by the salmon is roughly about See also:i000 to every See also:pound weight of the fish, a 15 lb salmon laying 15,000 eggs . The See also:sturgeon lays about 7,000,000; the See also:herring 5o,000; the turbot 14,311,000; the See also:sole 134,000; the perch 280,000 . Briefly, the number is greatest where the risks of destruction are greatest . The eggs of the degenerate fishes known as the lampreys and See also:hag-fishes are remarkable for the fact that in the latter they are large in size, cylindrical in shape, and provided at each end with hooklets whereby they adhere one to another; while in the lampreys they are extremely small and embedded in a jelly . Molluscs.—Among the See also:Mollusca, See also:Crustacea and Insecta yolk-stored eggs of very remarkable forms are commonly produced . In variety, in this connexion, the Mollusca must perhaps be given the first place . This diversity, indeed, is strikingly illustrated by the eggs of the See also:Cephalopoda . In the squids (Loligo), for example, the eggs are enclosed in long cylindrical cases, of which there are several hundreds, attached by one end to a common centre; the whole See also:series looking strangely like a rough See also:mop-See also:head . Each case, in such a cluster, contains about 250 eggs, or about 40,000 in all . By way of contrast the eggs of the true cuttle-fish (See also:Sepia) are deposited separately, each enclosed in a tough, black, pear-shaped capsule which is fastened by a stalk to fronds of sea-See also:weed or other See also:object . They appear to be extruded at See also:short intervals, till the full complement is laid, the whole forming a cluster looking like a bunch of grapes . The See also:octopus differs yet again in this matter, its eggs being very small, See also:berry-like, and attached to a stalk which runs through the centre of the mass . The eggs of the univalve Mollusca are hardly less varied in the shapes they take . In the common See also:British See also:Purpura lapillus they resemble delicate See also:pink grains of See also:rice set on stalks; in Busycon they are disk-shaped, and attached to a band nearly 3 ft. long .
The eggs of the shell-bearing slugs (Testacella) are large, and have the outer coat so elastic that if dropped on a See also: They are generally laid in clusters, either on the ground, on the leaves of plants, or in the water . Some of the gnats (Culex) lay them on the water . Cylindrical in shape they are packed closely together , set on end, the whole mass forming a kind of floating raft . Frequently, as in the case of the stick and leaf See also:insect, the eggs are enclosed in capsules of very elaborate shapes and highly ornamented . As to the rest of the Invertebrata—above the See also:Protozoa the eggs are laid in water, or in See also:damp places . In the former case they are as a rule small, and give rise to larvae; while eggs hatched on See also:land are sometimes enclosed in capsules, " cocoons," as in the case of the See also:earthworm, where this capsule is filled with a milky white fluid, of a highly nutritious character, on which the embryos feed . Among some invertebrates two different kinds of eggs are laid by the same individual . The water-See also:flea, Daphnia (a crustacean), lays two kinds of eggs known as " summer " and " See also:winter " eggs . The summer eggs are carried by the female in a " brood-pooch " on the back . The " winter " eggs, produced at the approach of winter, differ markedly in appearance from the summer eggs, being larger, darker in colour, thicker shelled, and enclosed in a capsule formed from the shell or See also:carapace, of the parent's body . " Winter eggs," however, may be produced in the height of summer . While the" summer eggs "are unfertilized, the winter eggs are fertilized by.the male, and possess the remarkable See also:power of lying dormant for months or even years before they develop . The See also:production of these two kinds of eggs is a device to overcome the See also:cold of winter, or the drying up of the pools in which the species lives, during the See also:heat of the summer . The power of resistance which such eggs possess may be seen in the fact that a See also:sample of mud which had been kept dry for ten years still contained living eggs . In deep water where neither drought nor winter cold can seriously affect the Daphnias, they propagate all the year See also:round by unfertilized " summer " eggs . Mollusca," Camb . Nat . Hist . (1906) . For further references to the above and other Invertebrate groups see various See also:text-books on See also:Entomology, See also:Zoology . (W . P . |
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