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ENTOMOLOGY (Gr. iv-roµa, insects, and...

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Originally appearing in Volume V09, Page 656 of the 1911 Encyclopedia Britannica.
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ENTOMOLOGY (Gr. iv-roµa, See also:insects, and Xiryos, a discourse)  , the See also:science that treats of See also:insects, i.e. of the animals included in the class See also:Hexapoda of the See also:great phylum (or sub-phylum) See also:Arthropoda . The See also:term, however, is somewhat elastic in its current use, and students of centipedes and See also:spiders are often reckoned among the entomologists . As the number of See also:species of insects is believed to exceed that of all other animals taken together, it is no wonder that their study should See also:form a See also:special See also:division of See also:zoology with a distinctive name . Beetles (Scarabaei) are the subjects of some of the See also:oldest sculptured See also:works of the Egyptians, and references to locusts, bees and ants are See also:familiar to all readers of the See also:Hebrew scriptures . The See also:interest of insects to the eastern races was, however,economic, religious or moral . The science of insects began with See also:Aristotle, who included in a class " Entoma " the true insects, the arachnids and the myriapods, the See also:Crustacea forming another class (" See also:Malacostraca") of the "Anaema" or " bloodless animals." For nearly 2000 years the few writers who dealt with zoological subjects followed Aristotle's leading . In the See also:history of the science, various lines of progress have to be traced . While some observers have studied in detail the structure and See also:life-history of a few selected types (See also:insect See also:anatomy and development), others have made a more superficial examination of large See also:series of insects to classify them and determine their relationships (systematic See also:entomology), while others again have investigated the habits and life-relations of insects (insect bionomics) . During See also:recent years the study of fossil insects (palaeoentomology) has attracted much See also:attention . The See also:foundations of See also:modern entomology were laid by a series of wonderful See also:memoirs on anatomy and development published in the 17th and 18th centuries . Of these the most famous are M . See also:Malpighi's See also:treatise on the silkworm (1669) and J .

See also:

Swammerdam's Biblia naturae, issued in 1737, fifty years after its author's See also:death, and containing observations on the structure and life-history of a series of insect types . Aristotle and See also:Harvey (De generatione animalium, 1651) had considered the insect larva as a prematurely hatched embryo and the pupa as a second See also:egg . Swammerdam, however, showed the presence under the larval cuticle of the pupal structures . His only unfortunate contribution to entomology—indeed to zoology generally—was his theory of pre-formation, which taught the presence within the egg of a perfectly formed but See also:miniature adult . A See also:year before Malpighi's great See also:work appeared, another See also:Italian naturalist, F . Redi, had disproved by experiment the spontaneous See also:generation of maggots from putrid flesh, and had shown that they can only develop from the eggs of flies . Meanwhile the See also:English naturalist, See also:John See also:Ray, was studying the See also:classification of animals ; he published, in 1705, his Methodus insectorum, in which the nature of the See also:metamorphosis received due See also:weight . Ray's " Insects " comprised the Arachnids, Crustacea, See also:Myriapoda and See also:Annelida, in addition to the . Hexapods . Ray was the first to formulate that definite conception of the species which was adopted by See also:Linnaeus and emphasized by his binominal nomenclature . In 1735 appeared the first edition of the Systema naturae of Linnaeus, in which the " Insecta " forma See also:group ' See also:equivalent to the Arthropoda of modern zoologists, and are divided into seven orders, whose names—See also:Coleoptera, See also:Diptera, See also:Lepidoptera, &c., founded on the nature of the wings—have become firmly established . The fascinating subjects of insect bionomics and life-history were dealt with in the classical memoirs (1734–1742) of the Frenchman R .

A . F. de See also:

Reaumur, and (1752–1778) of the Swede C. de Geer . The freshness, the See also:air of leisure, the See also:enthusiasm of See also:discovery that See also:mark the work of these old writers have lessons for the modern professional zoologist, who at times feels burdened with the accumulated knowledge of a See also:century and a See also:half . From the end of the 18th century until the See also:present See also:day, it is only possible to enumerate the outstanding features in the progress of entomology . In the See also:realm of classification, the work of Linnaeus was continued in See also:Denmark by J . C . See also:Fabricius (Systema entomologica, 1795), and extended in See also:France by G . P . B . See also:Lamarck (Animaux sans vertebres, 1801) and G . See also:Cuvier (Lecons d'anatomie comparee, 1800-1805), and in See also:England by W . E .

Leach (Trans . Linn . See also:

Soc. xi., 1815) . These three authors definitely separated the See also:Arachnida, Crustacea and Myriapoda as classes distinct from the Insecta (see HEXAPODA) . The work of J . O . Westwood (Modern Classification of Insects, 1839–1840) connects these older writers with their successors of to-day . In the anatomical See also:field the work of Malpighi and Swammerdam was at first continued most energetically by See also:French students . P . Lyonnet had published in 176o his elaborate monograph on the See also:goat-See also:moth See also:caterpillar, and H . E . See also:Strauss-Durckheim in 1828 issued his great treatise on the cockchafer .

But the name of J . C . L. de See also:

Savigny, who (Mem. sur See also:les animaux sans vertebres, 1816) established the homology of the jaws of all insects whether biting or sucking, deserves especial See also:honour . Many anatomical and developmental details were carefully worked out by L . See also:Dufour (in a See also:long series of memoirs from 1811 to z86o) in France, by G . See also:Newport (" Insecta " in Encyc . Anat. and Physiol., 1839) in England, and by H . Burmeister (Handbuch der Entomologie, 1832) in See also:Germany . Through the 19th century, as knowledge increased, the work of investigation became necessarily more and more specialized . Anatomists like F . Leydig, F . See also:Muller, B .

T . Lowne and V . Graber turned their attention to the detailed investigation of some one species or to special points in the structure of some particular See also:

organs, using for the elucidation of their subject the ever-improving microscopical methods of See also:research . See also:Societies for the discussion and publication of papers on entomology were naturally established as the number of students increased . The Societe Entomologique de France was founded in 1832, the Entomological Society of See also:London in 1834 . Few branches of zoology have been more valuable as a See also:meeting-ground for professional and See also:amateur naturalists than entomology, and not seldom has the amateur—as in the See also:case of Westwood—See also:developed into a See also:professor . During the pre-Linnaean See also:period, the beauty of insects—especially the Lepidoptera—had attracted a number of collectors; and these " Aurelians "—regarded as harmless lunatics by most of their See also:friends—were the forerunners of the systematic students of later times . While the insect See also:fauna of See also:European countries was investigated by See also:local naturalists, the spread of See also:geographical exploration brought ever-increasing stores of See also:exotic material to the great museums, and specialization —either in the fauna of a small See also:district or in the See also:world-wide study of an See also:order or a group of families—became constantly more marked in systematic work . As examples may be instanced the studies of A . H . Haliday and H . Loew on the European Diptera, of John See also:Curtis on See also:British insects, of H .

Phoenix-squares

T . Stainton and O . Staudinger on the European Lepidoptera, of R . M`Lachlan on the European and of H . A . See also:

Hagen on the See also:North See also:American See also:Neuroptera, of D . See also:Sharp on the Dyticidae and other families of Coleoptera of the whole world . The See also:embryology of insects is entirely a study of the last century . C . See also:Bonnet indeed observed in 1745 the virgin-See also:reproduction of Aphids, but it was not until 1842 that R . A. von See also:Kolliker described the formation of the blastoderm in the egg of the midge Chironomus . Later A .

See also:

Weismann (1863–1864) traced details of the growth of embryo and of pupa among the Diptera, and A . See also:Kovalevsky in 1871 first described the formation of the germinal layers in insects . Most of the recent work on the embryology of insects has been done in Germany or the See also:United States, and among numerous students V . Graber, K . Heider, W . M . See also:Wheeler and R . Heymons may be especially mentioned . The work of de Reaumur and de Geer on the bionomics and life-history of insects has been continued by numerous observers, among whom may be especially mentioned in France J . H . See also:Fabre and C . See also:Janet, in England W .

See also:

Kirby and W . See also:Spence, J . Lubbock (See also:Lord See also:Avebury) and L . C . See also:Miall, and in the United States C . V . See also:Riley . The last-named may be considered the founder of the strong See also:company of entomological workers now labouring in See also:America . Though Riley was especially interested in the See also:bearings of insect life on See also:agriculture and See also:industry—economic entomology (q. v.)—he and his followers have laid the science generally under a deep See also:obligation by their researches . After the publication of C . See also:Darwin's Origin of Species (18J9) a fresh impetus was given to entomology as to all branches of zoology, and it became generally recognized that insects form a group convenient and hopeful for the elucidation of certain problems of See also:animal See also:evolution . The writings of Darwin himself and of A .

R . See also:

Wallace (both at one See also:time active entomological collectors) contain much See also:evidence See also:drawn from insects in favour of descent with modification . The phylogeny of insects has since been discussed by F . Brauer, A . S . Packard and many others; See also:mimicry and allied problems by H . W . See also:Bates, F . Muller, E . B . Poulton and M . C .

Piepers; the bearing of insect habits on theories of selection and use-See also:

inheritance by A . Weismann, G . W. and E . See also:Peckham, G . H . T . Eimer and See also:Herbert See also:Spencer; variation by W . See also:Bateson and M . Standfuss . B1sL1oGRArxv.—References to the works of the above authors, and to many others, will be found under HExAronA and the special articles on various insect orders . Valuable summaries of the labours of Malpighi, Swammerdam and other See also:early entomologists are given in L . C .

Miall and A . Denny's Cockroach (London, 1886), and L . Henneguy's Les Insectes (See also:

Paris, 1904) . (G . H .

End of Article: ENTOMOLOGY (Gr. iv-roµa, insects, and Xiryos, a discourse)
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