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MOSQUITO (Span. mosquito, a gnat, dim...

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Originally appearing in Volume V18, Page 902 of the 1911 Encyclopedia Britannica.
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MOSQUITO (Span. mosquito, a See also:gnat, diminutive of mosca, a See also:fly)  , a See also:term originally applied to many See also:species of small See also:blood- A, Larva of Anopheles . F, See also:Female Anopheles costalis, B, Pupa of Anopheles . Loew . C, Larva of Stegomyia . G, See also:Head of 9 Culex . D, Pupa of Culex . H, Head of ' Anopheles . E, See also:Egg-See also:float and further enlarged J, Head ofd Culex . detached egg of Culex . K, Eggs of Anopheles . sucking See also:DIPTERA (q.v.), belonging to various families, but now by See also:common consent restricted to those known to naturalists as Culicidae, or gnats . Before the See also:year 1899 mosquitoes had never been collected systematically, and had received little See also:notice from entomologists, so that but few genera and comparatively few species were known .

Although it had See also:

long been suspected that these See also:insects were in some way connected with See also:malaria and other diseases, while that the species now called Stegomyia calopus was the See also:carrier of yellow See also:fever had been asserted by See also:Finlay as See also:early as 1881, it was not until the closing years of the 19th See also:century that the brilliant researches of See also:Ross in See also:India, and of Grassi and others in See also:Italy, directed the See also:attention of the whole civilized See also:world to mosquitoes as the exclusive agents in the dissemination of malarial fever . The result has been that in subsequent years mosquitoes have been collected, studied and described by naturalists and medical men in all parts of the globe . Nearly too genera and about 700 species of mosquitoes are now recognized, but in all See also:probability the See also:total number of species is not less than moo . In See also:general See also:appearance mosquitoes resemble many harmless midges (Chironomidae), but may be distinguished by the following characters . (r) The prolongation of the See also:lower See also:lip or labium into a prominent See also:proboscis, which in the female See also:sex contains the full See also:complement of piercing See also:organs found in blood-sucking Diptera, namely paired mandibles, paired maxillae, a tubular hypopharynx (the common outlet of the salivary glands), and an upper lip or See also:labrum . (2) The presence of variously formed scales on the See also:body and its appendages: the head is clothed with scales, the See also:thorax with hairs or scales, and the See also:abdomen with either hairs or scales, or both; the legs and See also:veins of the wings are always covered with scales, and the palpi are often (as in some Anophelinae) conspicuously scaly . (3) The fact that the costal or marginal vein runs completely See also:round the wing . The wings exhibit six See also:longitudinal veins (seven in Heptaphlebomyia), two of which are characteristically forked . The antennae, usually See also:bottle-See also:brush shaped (plumose) in the male sex, are less hairy in the female . The palpi vary in See also:form and in the number of their component segments, and the proboscis, though usually straight, may be curved (as in Megarhinus) or otherwise modified in shape . In dividing the Culicidae into genera reliance is placed chiefly , upon characters derived from the scales on the three divisions of the body and on the wings . A fairly satisfactory See also:attempt at grouping the genera has been made by Lutz (19o4), who divides the See also:family in the first See also:place into the Euculicidae, with a piercing proboscis (i.e. all See also:ordinary mosquitoes), and the Culicimorphae or forms with-out a piercing proboscis (Mochlonyx, Corethra, &c.) .

It has since been proposed to treat the Culicimorphae as a distinct family under the See also:

title Corethridae, and it is probable that with this modification Lutz's See also:scheme will meet with general See also:acceptance . The Euculicidae are divided into the Asiphonatae (=Anophelinae), the larvae of which have no See also:respiratory See also:siphon, and the Siphonatae, or forrns in which a respiratory siphon is See also:present in the larval See also:state . The divisions of the Siphonatae are the Ankylorhynchae (genera with curved proboscis, e.g . Megarhinus and Toxorhynchites) and Orthorhynchae (genera with straight proboscis) . The latter again are divided into Metanopsilae (in which the metanotum or posterior region of the thorax is See also:bare) and Metanotrichae (in which the metanoturn is clothed with bristles or scales) . The Metanopsilae are made up of the Heteropalpae [palpi long in the male, See also:short in the female; sub-families Culicinae (Culex, &c.) and Heptaphlebornyinae (Heptaphlebomyia)l and Micropalpae [palpi short in both sexes; sub-families Aedinae (Aedes, &c.) and Haemagoginae (Haemagogus, Uranotaenia, &c.)l . The Metanotrichae are similarly divided on the basis of the palpal characters into two See also:groups, the Heteropalpae or Hyloconopinae (Joblotia, Rhynchomyia, &c.) and Micropalpae or Dendromyinae (Wyeomyia, Sabethes, Limatus, &c.) . The old genus Anopheles (characterized by the palpi being long in both sexes) is now divided into a number of genera according to the See also:character and shape of the scales on the different regions of the body and on the wings . These genera make up the sub-family Anophelinae, and together include over too species . The genus Culex, from which the family takes its name, though it has been similarly split up, is still in its restricted sense larger than any other, and some 200 species are comprised in it alone . Mosquitoes are found in all parts of the world . Even within the See also:Arctic Circle they are in many localities abundant and excessively bloodthirsty during the short summer .

Phoenix-squares

Under such conditions the. deeply-rooted nature of the blood-sucking See also:

instinct is most remarkable; for insects whose ancestors for many generations may not have tasted blood will seek for it with the utmost keenness and pertinacity so soon as an opportunity presents itself . Some species are normally phytophagous, and the vast See also:majority, at any See also:rate, appear to be capable of continuing to exist and reproducing their See also:kind upon a purely vegetarian See also:diet . As a See also:rule the blood-sucking See also:habit is confined to the See also:females, but in the See also:case of a few species it is said to be common to both sexes . The thirst for blood is stimulated by See also:heat, and in temperate climates it is only during hot See also:weather that mosquitoes are troublesome . Some species of mosquitoes, such as the common See also:gnat (Culex pipiens), are rarely found away from human habitations; others seldom or never enter houses, but are met with either in more or less open See also:country, or in the recesses of forests and See also:woods . In See also:Europe and See also:North See also:America the continued existence of species is ensured by the See also:hibernation of impregnated females, or else the See also:winter is passed in the egg or occasionally in the larval state . In tropical climates with a well-marked dry See also:season mosquitoes pass into a semi-dormant See also:condition during the See also:period when there is little See also:water in which to See also:deposit their eggs . Culicidae are by no means confined to See also:low-lying districts, and have even been met with in the Himalayas at an See also:altitude of 13,000 feet . The wide See also:distribution of certain species is undoubtedly attributable to the agency of See also:ships and trains; under natural conditions mosquitoes seldom travel far from their breeding grounds,. although the See also:powers of See also:flight of some species are greater than has been supposed . The preliminary stages of all mosquitoes are passed in water, either fresh or See also:salt, stagnant or slightly moving . The nature of the breeding-place varies greatly according to the species, and while many of the mosquitoes that infest houses will breed even in the smallest accidental See also:accumulation of water such as may have collected in a discarded bottle or See also:tin, the larvae of other species less closely associated with See also:man are found in natural pools or ditches, at the margins of slow-moving streams, in collections of water in hollow trees and See also:bamboo-stumps, or even in the water-receptacles of certain See also:plants . The eggs are usually deposited on the water itself, and while in the case of certain species, such as Culex pipiens or the widely distributed C. fatigans, they are agglutinated together in masses known as " boats " or " rafts " containing from 50 to 400 ova, those of others, such as the Anophelinae and many Culicinae (e.g .

Stegomyia calopus), are laid separately . The larvae are active and voracious little See also:

grub-like creatures (known in the See also:United States as " wrigglers "), with large heads and jaws provided with a pair of brushes, which sweep See also:food-particles into the mouth . Their food consists of See also:minute See also:animal and See also:vegetable organisms, (Redrawn by permission from Farmers Bulletin 155, See also:Bureau of Ent., U.S . Dept. of See also:Agriculture.) See also:algae, and probably decaying vegetable See also:matter; they are often cannibals, and feed on their own species . The larvae of species belonging to the Culicinae have a prominent breathing See also:tube, or respiratory siphon, on the penultimate (eighth) abdominal segment, and when taking in See also:air hang head downwards (often nearly vertically) from the See also:surface film . Larvae of Anophelinae, on the other See also:hand—which are See also:grey, See also:green or See also:brown in See also:colour, and often extremely difficult to see—have no respiratory siphon and See also:lie almost horizontally at the surface; they frequently appear as though anchored by the tail to a See also:weed or other See also:object, and possess the curious See also:faculty of completely rotating the head so as to browse on the surface film . See also:Mosquito pupae are See also:comma-shaped (see fig . I), and breathe by means of a pair of respiratory trumpets on the thorax . The majority of mosquitoes are dull in See also:hue, but certain species are brilliantly coloured or conspicuously banded or spotted with See also:white . The Anophelinae have narrow bodies, and generally spotted wings, and when at See also:rest keep body and proboscis in a straight See also:line, often at a considerable See also:angle with the supporting surface; in this way they can be distinguished from Culicinae, which have a humped-up thorax with which the proboscis forms an angle, and in the resting position keep the body parallel to the support . The disseminators of malaria are exclusively Anophelinae, but even among these it is only certain species that are dangerous, since the others appear to be incapable of acting as hosts of the parasites . Stegomyia calopus, on the other hand, a very widely distributed species and the almost certain carrier of yellow fever, belongs to the Culicinae .

In the case of See also:

filariasis due to Filaria bancrofti, which is common throughout the Tropics, the embryos of the See also:parasite are disseminated by various Culicinae and Anophelinae (Culex pipiens in See also:Queensland; C. fatigans in the See also:West Indies; Myzomyia rossii in India; Pyretophorus costalis in a large portion of tropical See also:Africa; &c.) . Six or seven species of mosquitoes are also the intermediate hosts of Filaria immitis, which infests the right See also:auricle and pulmonary artery of the See also:dog, and occurs throughout the tropics, in See also:southern Europe, the United States of America, and elsewhere . There is See also:reason to believe that malaria, yellow fever and filariasis are not the only diseases disseminated by mosquitoes . (E . E .

End of Article: MOSQUITO (Span. mosquito, a gnat, diminutive of mosca, a fly)
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