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MYLODON (Gr. for " See also: extinct See also: American edentate mammals, typified by a See also: species (M. harlani) from the See also: Pleistocene of See also: Kentucky and other parts of the See also: United States, but more abundantly represented in the corresponding formations of See also: South See also: America, especially See also: Argentina and See also: Brazil
.
The mylodons belong to the See also: group of ground-sloths, and are generally included in the See also: family Megatheriidae, although sometimes made the type of a See also: separate family
.
From Megatherium these animals, which rivalled the Indianrhinoceros in bulk, differ in the shape of their cheek-teeth; these (five above and four below) being much smaller, with an ovate section, and a cupped instead of a ridged See also: crown-See also: surface, thus resembling those of the true sloths
.
In certain species of mylodon the front pair of teeth in each jaw is placed some distance in front of the rest and has the crown surface obliquely bevelled by,
From See also: Owen
.
See also: Skeleton of Mylodon robustus (Pleistocene, South America)
.
wearing against the corresponding teeth in the opposite jaw
.
On this account such species have been referred to a second genus, under the name of Lestodon, but the distinction scarcely seems necessary
.
The See also: skull is shorter and See also: lower than in Megatherium, without any vertical expansion of the See also: middle of the lower jaw, and the teeth also extend nearly to the front of the jaws; both these features being See also: sloth-like
.
In the fore feet the three inner toes have large claws, while the two See also: outer ones are rudimentary and clawless; in the See also: hind-limbs the first toe is wanting, as in Megatherium, but the second and third are clawed
.
The skin was strengthened by a number of small deeply-embedded bony nodules
.
Although the typical M. harlani is North—American, the mylodons are essentially a South American group, a few of the representatives of which effected an entrance into See also: North America when that continent became finally connected with South America
.
See also: Special See also: interest attaches to the See also: recent See also: discovery in the cavern of Ultima Esperanza, South See also: Patagonia, of remains of the genus Glossotherium, or Grypotherium, a near relative of Mylodon, but differing from it in having a bony See also: arch connecting the nasal bones of the skull with the premaxillae; these include a considerable portion of the skin with the hair attached
.
Ossicles somewhat resembling large See also: coffee-berries had been previously found in association with the bones of Mylodon, and in Glossotherium nearly similar ossicles occur embedded on the inner See also: side of the thick hide
.
The coarse and shaggy hair is somewhat like that of the sloths
.
The remains, which include not only the skeleton and skin, but likewise the droppings, were found buried in grass which appears to have been chopped up by See also: man, and it thus seems not only evident that these ground-sloths dwelt in the cave, but that there is a considerable probability of their having been kept there in a semi-domesticated See also: state by the early human inhabitants of Patagonia
.
The extremely fresh condition of the remains has given rise to the idea that Glossotherium may still be living in the wilds of Patagonia
.
Scelidotherium is another genus of large South American Pleistocene ground-sloths, characterized, among other features, by the See also: elongation and slenderness of the skull, which thus makes a decided approximation to the See also: anteater type, although retaining the full series of cheek-teeth, which were, of course, essential to an herbivorous animal
.
The feet resemble those of Megatherium
.
A much smaller South American species represents the genus Nothrotherium
.
In North America Mylodon was accompanied by another gigantic species typifying the genus Megalonyx, in which the fore See also: part of the skull was usually wide, and the third and See also: fourth front toes carried claws
.
Another genus has been described from the Pleistocene
of See also: Nebraska, as Paramylodon; it has only four pairs of teeth, and an elongate skull with an inflated muzzle
.
All the above genera differ from Megatherium in having a foramen on the inner side of the lower end of the humerus
.
A presumed large ground-sloth from See also: Madagascar has been described, on the evidence of a See also: limb-See also: bone, as Bradytherium, but it is suggested by Dr F
.
Ameghino that the specimen really belongs to a lemuroid
.
Be this as it may, the North American mammals described as See also: Moro pus and Morotherium, in the belief that they were ground-sloths, are really referable to the ungulate group Ancvlopoda
.
Although a few of the Pleistocene ground-sloths, such as Nothropus and Nothrotherium (= Coelodon), were of comparatively small See also: size, in the See also: Santa Cruz beds of Patagonia few of the representatives of the family much exceeded a See also: modern sloth in size
.
The best-known generic types are Eucholoeops, Hapalops and Pseudahapalops, of which considerable portions of the skeleton have been disinterred
.
In these diminutive ground-sloths the crowns of the cheek-teeth approached the prismatic See also: form characteristic of Mega[lo]therium, as distinct from the subcylindrical type occurring in Mylodon, Glossotherium, &c
.
By many palaeontologists a group of North American Lower See also: Tertiary mammals, known as Ganodonta, has been regarded as representing the ancestral stock of the ground-sloths and those of other South American edentates; but according to Professor W
.
B
.
See also: Scott this view is incorrect and there is no See also: affinity between the two See also: groups
.
If this be so, we are still in See also: complete darkness as to the stock from which the South American edentates are derived
.
See W
.
B
.
Scott, Mammalia of the Santa Cruz Beds, See also: Edentata, See also: Rep., See also: Princeton Exped. to Patagonia, vol. v
.
(1903—1904); B
.
See also: Brown A New Genus of Ground-Sloth from the Pleistocene of Nebraska, Bull
.
Amer
.
See also: Mus
.
Nat
.
Hist., xix, 569 (1903)
.
(R
.
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