Online Encyclopedia

Search over 40,000 articles from the original, classic Encyclopedia Britannica, 11th Edition.

ORDER I

Online Encyclopedia
Originally appearing in Volume V26, Page 543 of the 1911 Encyclopedia Britannica.
Spread the word: del.icio.us del.icio.us it!

See also:

ORDER I  .—CROSSOPTERYGII Paired fins, at least the pectorals, lobate, having an endo-skeletal See also:axis more or less fringed with dermal rays . Mandibular See also:arch suspended from the upper segment of the hyoid arch (hyostylic See also:skull) . Splenial See also:bone See also:present . No supraoccipital bone . A pair of large jugular plates, sometimes with small lateral plates and an anterior azygous See also:element, See also:developed in the branchiostegal membrane between the mandibular rami . See also:Heart with a' contractile, multivalvular conus arteriosus; See also:intestine with a See also:spiral See also:valve; See also:air-See also:bladder with pneumatic duct communicating with the ventral See also:side of the See also:oesophagus . Maxillary bone large, toothed, bordering the mouth . Bones of the upper See also:surface of the skull mostly paired . See also:Pectoral arch with both clavicle (so-called infra-clavicle) and cleithrum . Ventral fins inserted far back . With few exceptions (tail of Coelacanthidae, dorsal and caudal fins of Polypteridae) the dermal rays of the unpaired fins more numerous than their endo-skeletal supports, a See also:primitive See also:character also found in the See also:lower Ganoids, but disappearing in the higher . SUB-See also:ORDER I.—OSTEOLEPIDA (Including the Haplistia, Rhipidistia and Actinistia.) Pectoral fins obtusely or acutely lobate, articulating with the pectoral See also:girdle by a single basal endo-skeletal element .

Nostrils on the lower side of the snout . Two dorsal fins . Families: Osteolepidae, Rhizodontidae, Holoptychidae, Coelacanthidae . The scales may be rhombic and thickly coated with ganoine (Osteolepidae) or See also:

cycloid . The vertebral axis is strongly heterocercal in the Osteolepidae and Holoptychidae, and diphycercal or intermediate between the heterocercal and the diphycercal types in the other families; usually acentrous, sometimes with See also:ring-like calcifications (some of the Rhizodontidae) . In the Holoptychidae the pectoral fin is extremely similar to that of the Dipneusti of the See also:family Dipteridae, which they resemble closely in See also:form and scaling . Their See also:teeth are remarkable for their complicated structure, resembling that of the Labyrinthodont Batrachians . A pineal foramen is present between the frontal bones in most of the Rhizodontidae . The Osteolepidae were mostly moderate-sized fishes, the largest (Megalichthys) measuring about 4 ft. in length . These Crossopterygians first appear in the Lower Devonian, are abundant in the Upper Devonian, Carboniferous and See also:Permian; in later periods they are represented only by the more specialized Coelacanthidae, which appear in the Lower Carboniferous, and persist as See also:late as the Upper See also:Chalk . SUB-ORDER II.—CLADISTIA Pectoral fin obtusely lobate, with three basal endo-skeletal elements . Nostrils on the upper side of the snout .

A single dorsal fin, formed of a See also:

series of detached rays . A single family: Polypteridae . The existing Crossopterygians which form this sub-order differ very considerably from the See also:extinct Osteolepida, perhaps quite as much as these differ from the Dipneusti . The ventral fins are not lobate, the vertebral See also:column is well ossified and its termination is of the diphycercal type . Spiracles, covered by bony valves, are present on the upper surface of the See also:head . The dorsal fin is unique among fishes, being formed of detached rays consisting of a spine-like fulcra! See also:scale supporting the fringes of the See also:ray; these rays have been regarded, erroneously, as representing so many distinct fins, or " finlets." The scales are bony, rhombic and thickly coated with ganoine . The Polypteridae are confined to tropical See also:Africa and the See also:Nile, and represented by two genera: Polypterus and Calamichthys, the former moderately elongate and provided with ventral fins, the latter serpentiform and devoid of ventrals . We now know ten See also:species of Polypterus, from the Nile, the See also:Congo, the See also:rivers of See also:West Africa, and lakes See also:Chad, See also:Rudolf and See also:Tanganyika, and one of Calamichthys, which inhabits West Africa from the See also:Niger See also:delta to the Chiloango . The largest species of Polypterus reach a length of nearly 4 ft . The See also:young are provided with an See also:external opercular gill very similar to the gills of larval salamanders . The air-bladder acts as an See also:accessory breathing See also:organ, although these fishes are not known ever to leave the See also:water . The development is stated by the late J .

S . Budgett to be even more Batrachian-like than that of the Dipneusti, but the results of the study of the material collected by him shortly before his See also:

death have not yet been published .

End of Article: ORDER I
[back]
ORDER 3
[next]
ORDER II

Additional information and Comments

There are no comments yet for this article.
» Add information or comments to this article.
Please link directly to this article:
Highlight the code below, right click and select "copy." Paste it into a website, email, or other HTML document.