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ROOT (late O.E. rot, adopted from Sca...

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Originally appearing in Volume V23, Page 713 of the 1911 Encyclopedia Britannica.
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ROOT (See also:late O.E. rot, adopted from Scand., cf. Norw. and Swed. rot, See also:Dan. See also:rod; the true O.E. word was wyrt, plant, represented in Ger. Wurz or Wurzel; the ultimate root is the same in both words, and is seen in See also:Lat. radix)  , the underground See also:part of a plant . This is the popular meaning of the word . In its botanical use the See also:term is more restricted (see below) . The various other meanings have all See also:developed from this, its See also:primary, significance . Of these the See also:principal are: the source or origin of a See also:condition, See also:state, quality, &c.; the See also:base or embedded part of a structure of the See also:body, such as a See also:nail, tooth, the See also:hair, &c.; in See also:mathematics, a number, quantity or See also:dimension which produces a given expression when multiplied by itself a requisite number of times; and in See also:philology an ultimate See also:element of See also:language, incapable of further See also:analysis . A particular See also:extension of the primary meaning is that which applies the word generally to a class of See also:plants, such as the See also:turnip or See also:carrot, whose See also:root is fleshy, and edible either by See also:man or domestic animals . The embryo of a. typical plant, for instance a See also:pea plant (fig . I), has an ascending See also:axis which will grow into the shoot, and a descending axis or radicle which will grow into the root. y When the See also:seed germinates, the radicle is the first to appear; it grows downwards, and its primary See also:function is to See also:act as a holdfast for the plant; its most important function, however, is the absorption of See also:water and . r.—The Dicotyledonous dissolved nutrient substances FIC Embryo of the Pea laid open. from the See also:soil, and it also frec, c, the two fleshy coty- quently serves for storage of foodledons, or seed-lobes, which stuffs . The root is distinguished remain under ground when from underground shoots by not the plant sprouts; r, the bearing leaves and by having radicular extremity of the axis which develops into its See also:apex (growing point) See also:pro-the root; 1, the axis bearing tected by a cap (root-cap), which the See also:young stalk and leaves can be clearly seen by makg, which See also:lie in a depression See also:ing a median See also:vertical See also:section of the cotyledons f . through the root-tip; the cap protects it in its passage through the soil . The root also generally bears root-hairs, slender unicellular outgrowths of the See also:outer layer, See also:borne in the region a little behind the root-tip . It is by means of the root - hairs especially that the root is brought into See also:close relation with the soil particles and absorbs the nutrient materials in See also:solution in the water which surrounds these particles .

The older root-hairs are continually dying off, so that they are borne only on a small part of the See also:

area behind the apex . Branches of the root, which repeat the See also:form and structure of the See also:main root, are developed in See also:regular See also:succession from above downwards (acropetal), and owing to the fact that they originate in a definite position in the interior of the root (endogenous) they develop in See also:longitudinal rows and have to break through the overlying See also:tissue of the See also:parent root (fig . 2) . True forking of the root (See also:dichotomy) occurs in the Lycopodiaceae (the shoots of which also See also:branch dichotomously), but is unknown in the higher plants . Roots which originate elsewhere than as acropetal out. growths of a main root are known as See also:adventitious, and may From See also:Green's See also:Vegetable See also:Physiology, by permission . Root-hair in contact with See also:par- ( Ultimate root-branches, showing tides of soil (highly magnified). position of root-hairs . arise on any part of a plant . They are especially numerous on underground stems, such as the under See also:side of rhizomes, and also develop from See also:stem nodes under favourable conditions, such as moisture and See also:absence of See also:light; a young shoot or a cutting placed in moist soil quickly forms adventitious roots . They may also arise from leaves under similar conditions, as, for instance, from See also:begonia leaves when planted in soil . The forms of roots depend on their shape and mode of branching . When the central axis goes deep into the ground in a tapering manner, without dividing, a tap-root is produced . This See also:kind of root is some-times shortened, and becomes swollen by storage of See also:food-stuffs, forming the conical root of carrot, or the fusiform or spindle-shaped root of See also:radish, or the napiform root of turnip .

In See also:

ordinary See also:forest trees the first root protruded continues to elongate and forms a See also:long primary root-axis, whence secondary axes come off . In primary plants, especially Monocotyledons, the primary axis soon See also:dies and the secondary axes take its See also:place . When the descending axis is very See also:short, and at once divides into thin, nearly equal fibrils, the root is called fibrous, as in many See also:grasses (fig . 4) ; when the fibrils are thick and succulent, the root is fasciculated, as in See also:Ranunculus Ficaria, Asphodelus luteus, and 0enanthe crocata; when some of the fibrils are developed in the form of tubercules, the root is tubercular, as in See also:dahlia (fig . 5) ; when the fibrils enlarge in certain parts only, the root is-nodulose, as in Spiraea Filipendula, or moniliform, as in Pelargonium triste, or annulated, as in See also:Ipecacuanha . Some of these so-called roots are formed of a stem and root combined, as in Orchis (fig . 6), where the tuber consists of a fleshy swollen From Vines's Student's permission . See also:Botany, by -/ From Strasburger's Lehrbuch der Botanik, by per- See also:mission of Gustav See also:Fischer . root bearing at the apex a stem bud . As in the See also:case of the stem, growth in length occurs only for a short distance behind the apex, but in long-lived roots increase in See also:diameter occurs continually in a similar manner to growth in thickness in the stem . Roots are usually underground and See also:colour-less, but in some cases where they arise from the stem they pass for some distance through the See also:air before reaching the soil . Such roots are called aerial .

They are well seen in the See also:

screw-See also:pine (Pandanus), the See also:Banyan (Ficus indica, fig . 7), and many other See also:species of Ficus, where they assist in supporting the stem and branches . In the See also:mangrove they often form the entire support of the stem, which has decayed at its See also:lower part . In See also:tree-ferns they form a dense coating around, and completely concealing, the stem; such is also the case in some Dracaenas and palms . In . Epiphytes, or plants growing in the air, attached to the trunks of trees, such as See also:orchids of warm climates, the aerial roots produced Fin . 6.-Base of do not reach the soil; they continue always plant of On-his, aerial and greenish, and they possess stomata. showing tuber- Delicate hairs are often seen on these epicules or tuberous Phytal roots, as well as a See also:peculiar spongy roots. investment formed by the cells of the epi- dermis which have lost their succulent See also:con-tents and are now filled with air . This layer is called the velamen, and serves to condense the moisture contained in the air, on which aerial roots, which reach the soil, and prop the branches . the plant is dependent for its water-See also:supply . The aerial roots of the See also:ivy are not the nutritive roots of the plant, but are only intended for See also:mechanical support . The climbing roots of many orchids, aroids and epiphytic ferns branch and form places of lodgment for humus into which absorbent branches of the climbing roots penetrate . Some leafless epiphytic orchids, such as species of Angraecum, depend entirely upon their aerial roots for nourishment; the roots, which are green, perform the functions both of leaves and roots .

A See also:

respiratory or aerating function is performed by roots of certain mangroves growing in swampy soil or water and sending vertical roots up into the air which are provided with aerating passages by which the root See also:system below can communicate with true outside air . Parasitic plants, as the mistletoe (Viscum), See also:broom-See also:rape (Orobanche) and Raflesia, send root-like processes into the substance of the plants whence they derive nourishment . In the See also:dodder (Cuscuta) , the tissue around the root swells into a kind of sucker (haustorium), which is applied See also:flat upon the other plant, and ultimately becomes See also:concave, so as to attach the plant by a vacuum . From the bottom of the sucker the root protrudes, and penetrates the tissue of the See also:host plant . See also:Leaf-buds are sometimes formed on roots, as in See also:plum, See also:cherry and other See also:fruit trees; the See also:common See also:elm affords an excellent example, the young shoots which grow up in the neighbourhood of a tree arising from the roots beneath the soil . In some plants no roots are formed at all; thus in the orchid Corallorhiza, known as See also:coral-root, a stem-structure, the shortly branched underground rhizome, performs all the functions of a true root which is absent . In aquatic plants the root acts merely as a holdfast or is altogether absent as in Salvinia, Utricularia and others .

End of Article: ROOT (late O.E. rot, adopted from Scand., cf. Norw. and Swed. rot, Dan. rod; the true O.E. word was wyrt, plant, represented in Ger. Wurz or Wurzel; the ultimate root is the same in both words, and is seen in Lat. radix)
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THEODORE ROOSEVELT (1858— )
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ELIHU ROOT (1845– )

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