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See also:ZOOLOGY (from Gr. Nov, a living thing, and ?byos, theory) , that portion of See also:biology (q.v.) which relates to animals, as distinguished from that portion (See also:Botany) which is concerned with See also:plants . See also:HISTORY There is something almost pathetic in the childish wonder and delight with which mankind in its earlier phases of See also:civilization gathered up and treasured stories of See also:strange animals from distant lands or deep seas, such as are recorded in the See also:Physiologus, in Albertus See also:Magnus, and even at the See also:present See also:day in the popular See also:treatises of See also:Japan and See also:China . That omnivorous universally credulous See also:stage, which may be calledthe " legendary," was succeeded by the See also:age of collectors and travellers, when many of the strange stories believed in were actually demonstrated as true by the living or pre- See also:change served trophies brought to See also:Europe . The possibility of in the verification established verification as a See also:habit; and See also:character the See also:collecting of things, instead of the accumulating See also:ing of reports, See also:developed a new See also:faculty of See also:minute observation . The See also:early collectors of natural curiosities were the founders of zoological See also:science, and to this day the naturalist-traveller and his correlative, the museum See also:curator and systematist, See also:play a most important See also:part in the progress of See also:zoology . Indeed, the See also:historical and present importance of this aspect or See also:branch of zoological science is so See also:great that the name " zoo-logy " has until recently been associated entirely with it, to the exclusion of the study of minute anatomical structure and See also:function which have been distinguished as See also:anatomy and See also:physiology . Anatomy and the study of See also:animal mechanism, animal physics and animal See also:chemistry, all of which See also:form part of a true zoology, were excluded from the usual See also:definition of the word by the See also:mere See also:accident that the zoologist had his museum but not his See also:garden of living specimens as the botanist had;' and, whilst the zoologist was thus deprived of the means of anatomical and physiological study—only later supplied by the method of preserving animal bodies in See also:alcohol—the demands of See also:medicine for a knowledge of the structure of the human animal brought into existence a See also:separate and See also:special study of human anatomy and physiology . From these special studies of human structure the knowledge of the anatomy of animals has proceeded, the same investigator who had made himself acquainted with the structure of the human See also:body desiring to compare with the See also:standard given by human anatomy the structures of other animals . Thus See also:comparative anatomy came into existence as a branch of inquiry apart from zoology, and it was only in the latter 'part of the 19th See also:century that the See also:limitation of the word " zoology " to a know-ledge of animals which expressly excludes the See also:consideration of their See also:internal structure was rejected by the See also:general See also:con-sent of those concerned in the progress of science . It is now generally recognized that it is mere tautology to speak of zoology and comparative anatomy, and that museum naturalists must give See also:attention as well to the inside as to the outside of animals . Scientific zoology really started in the 16th century with the awakening of the new spirit of observation and exploration, but for a See also:long See also:time ran a separate course uninfluenced by the progress of the medical studies of anatomy and physiology . The active See also:search for knowledge by means of observation and experiment found its natural See also:home in the See also:universities .
Owing to the connexion of medicine with these seats of learning, it was natural that the study of the structure and functions of the human body and of the animals nearest to See also:man should take See also:root there; the spirit of inquiry which now for the first time became general showed itself in the anatomical See also:schools of the See also:Italian universities of the 16th century, and spread fifty years later to See also:Oxford
.
In the 17th century the lovers of the new See also:philosophy, the investigators of nature by means of observation and experiment, banded themselves into See also:academies or See also:societies for mutual support and intercourse
.
The first founded of surviving See also:European academies, the Academia Naturae Curiosorum (1651),2 especially confined itself to the description and See also:illustration of the structure of plants and animals; eleven years later (1662) the Royal Society of See also:London was incorporated by royal See also:charter, having existed without a name or fixed organization for
1 The See also:medieval attitude towards both plants and animals had no relation to real knowledge, but was part of a See also:peculiar and in itself highly interesting See also:mysticism
.
A fantastic and elaborate See also:doctrine of symbolism existed which comprised all nature; See also:witchcraft, See also:alchemy and medicine were its See also:practical expressions
.
Animals as well as plants were regarded as " simples " and used in medicine, and a knowledge of them was valued from this point of view
.
2 The Academia Secretorum Naturae was founded at See also:Naples in 156o, but was suppressed by the ecclesiastical authorities
.
seventeen years previously (from 1645)
.
A little later the See also:Academy of Sciences of See also:Paris was established by See also: The perfecting of the microscope led to a full comprehension of the great doctrine of See also:cell-structure and the See also:establishment of the facts—(r) that all organisms are either single corpuscles (so-called cells) of living material (microscopic animalcules, &c.) or are built up of an immense number of such See also:units; (2) that all organisms begin their individual existence as a single unit or corpuscle of living substance, which multiplies by binary fission, the products growing in See also:size and multiplying similarly by binary fission; and (3) that the See also:life of a multi-cellular organism is the sum of the activities of the corpuscular units of which it consists, and that the processes of life must be studied in and their explanation obtained from an under-See also:standing of the chemical and See also:physical changes which go on in each individual corpuscle or unit of living material or See also:protoplasm . Meanwhile the astronomical theories of development of the See also:solar See also:system from a gaseous See also:condition to its present form, put forward by See also:Kant and by See also:Laplace, had impressed men's minds ideas of with the conception of a general See also:movement of spondevelop- taneous progress or development in all nature . The `neat. science of See also:geology came into existence, and the whole See also:panorama of successive stages of the See also:earth's history, each with its distinct See also:population of strange animals and plants, unlike those of the present day and simpler in proportion as they recede into the past, was revealed by See also:Cuvier, See also:Agassiz and others . The history of the crust of the earth was explained by See also:Lyell as due to a See also:process of slow development, in See also:order to effect which he called in no cataclysmic agencies, no mysterious forces differing from those operating at the present day . Thus he carried on the narrative of orderly development from the point at which it was See also:left by Kant and Laplace—explaining by reference to the ascertained See also:laws of physics and chemistry the configuration of the earth, its mountains and seas, its igneous and its stratified rocks, just as the astronomers had explained by those same laws the See also:evolution of the sun and See also:planets from diffused gaseous See also:matter of high temperature . The See also:suggestion that living things must also be included in this great development was obvious . The delay in the establishment of the doctrine of organic evolution was due, not to the ignorant and unobservant, but to the leaders of zoological and botanical science . Knowing the almost endless complexity of organic structures, realizing that man himself with all the See also:mystery of his life and consciousness must be included in any explanation of the origin of living things, they preferred to regard living things as something apart fromthe See also:rest of nature, specially cared for, specially created by a Divine Being . Thus it was that the so-called " Natur-philosophen " of the last See also:decade of the 18th century, and The their successors in the first See also:quarter of the 19th, Nalurfound few adherents among the working zoologists tiro-and botanists . See also:Lamarck, Treviranus, See also:Erasmus See also:Dar- peen. win, See also:Goethe, and See also:Saint-Hilaire preached to See also:deaf ears, for they advanced the theory that living beings had developed by a slow process of transmutation in successive generations from simpler ancestors, and in the beginning from simplest formless matter, without being able to demonstrate any existing See also:mechanical causes by which such development must necessarily be brought about . They were met by the See also:criticism that possibly such a development had taken See also:place; but, as no one could show as a See also:simple fact of observation that it had taken place, nor as a result of legitimate inference that it must have taken place, it was quite as likely that the past and present See also:species of animals and plants had been separately created or individually brought into existence by unknown and inscrutable causes, and (it was held) the truly scientific man would refuse to occupy himself with such fancies, whilst ever continuing to concern himself with the observation and See also:record of indisputable facts . The critics did well; for the " Natur-philosophen," though right in their See also:main conception, were premature .
It was reserved for See also: It cannot be said that previously to Darwin there had been New dc- any very profound study of teleology, but it had vetopment been the delight of a certain type of mind—that of of teteo- the lovers of nature or naturalists See also:par excellence, as logy, they were sometimes termed—to See also:watch the habits of living animals and plants, and to point out the remarkable ways in which the structure of each variety of organic life was adapted to the special circumstances of life of the variety or species . The astonishing See also:colours and See also:grotesque forms of some animals and plants which the museum zoologists gravely de-scribed without comment were shown by these observers of living nature to have their significance in the See also:economy of the organism possessing them; and a general doctrine was re-cognized, to the effect that no part. or structure of an organism is without definite use and adaptation, being designed by the Creator for the benefit of the creature to which it belongs, or else for the benefit, amusement or instruction of his highest creature—man . Teleology in this form of the doctrine of See also:design was never very deeply rooted amongst scientific anatomists and systematists . It was considered permissible to speculate somewhat vaguely on the subject of the utility of this or that startling variety of structure; but few attempts, though some of great importance, were made systematically to explain by observation and experiment the adaptation of organic structures to particular purposes in the See also:case of the See also:lower animals and plants . Teleology had, indeed, an important part in the development of physiology—the knowledge of the mechanism, the physical and chemical properties, of the parts of the body of man and the higher animals allied to hint . But, as applied to lower and more obscure forms of life, teleology presented almost insurmountable difficulties; and consequently, in place of exact experiment and demonstration, the most reckless though ingenious assumptions were made as to the utility of the parts and See also:organs of lower animals . Darwin's theory had as one of its results the See also:reformation and rehabilitation of teleology . According to that theory, every See also:organ, every part, See also:colour and peculiarity of an organism, must either be of benefit to that organism itself or have been so to its ancestors:1 no peculiarity of structure or general conformation, no habit or See also:instinct in any organism, can be supposed to exist for the benefit or amusement of another organism, not even for the delectation of man him-self . Necessarily, according to the theory of natural selection, structures either are present because they are selected as useful or because they are still inherited from ancestors to whom they were useful, though no longer useful to the existing representatives of those ancestors . Structures previously inexplicable were now explained as survivals from a past age, no longer useful though once of value . Every variety of form and colour was urgently and absolutely called upon to produce its See also:title to existence either as an active useful See also:agent or as a survival . Darwin himself spent a large part of the later years of his life in thus extending the new teleology . The old doctrine of types, which was used by the philosophically minded zoologists (and botanists) of the first See also:half 1 A very subtle and important qualification of this generalization has to be recognized (and was recognized by Darwin) in the fact that owing to the interdependence of the parts of the bodies of living things and their profound chemical interactions and peculiar structural See also:balance (what is called organic See also:polarity) the variation of one single part (a spot of colour, a tooth, a claw, a leaflet) may, and demonstrably does in many cases See also:entail variation of other parts—what are called correlated variations . Hence many structures which are obvious to the See also:eye, and serve as distinguishing marks of separate species, are really not themselves of value or use, but are the necessary concomitants of less obvious and even altogether obscure qualities, which are the real characters upon which selection is acting . Such " correlated variations " may attain to great size and complexity without being of use . But eventually they may in turn become, in changed conditions, of selective value . Thus in many cases the difficulty of supposing that selection has acted on minute and imperceptible initial variations, so small as to have no selective value, may be got rid of . A useless " correlated variation " may have attained great See also:volume and quality before it is (as it were) seized upon and perfected by natural selection . All organisms are essentially and necessarily built up by such correlated variations.of the 19th century as a ready means of explaining the failures and difficulties of the doctrine of design, See also:fell into its proper place under the new See also:dispensation . The adherence to type, the favourite conception of the transcendental morphologist, was seen to be nothing more than the expression of one of the laws of thremmatology, the persistence of hereditary trans-See also:mission of ancestral characters, even when they have ceased to be significant or valuable in the struggle for existence, whilst the so-called evidences of design which was supposed to modify the limitations of types assigned to Himself by the Creator were seen to be adaptations due to the selection and intensification by selective breeding of fortuitous congenital variations, which happened to prove more useful than the many thousand other variations which did not survive in the struggle for existence . Thus not only did Darwin's theory give a new basis to the study of organic structure, but, whilst rendering the general theory of organic evolution equally acceptable and Effects or necessary, it explained the existence of See also:low and simple Darwin's forms of life as survivals of the earliest ancestry of theory more highly complex forms, and revealed the classi- "p0° fications of the systematist as unconscious attempts zoology. to construct the genealogical See also:tree or See also:pedigree of plants and animals . Finally, it brought the simplest living matter or formless protoplasm before the See also:mental See also:vision as the starting-point whence, by the operation of necessary mechanical causes, the highest forms have been evolved, and it rendered unavoidable the conclusion that this earliest living material was itself evolved by See also:gradual processes, the result also of the known and recognized laws of physics and chemistry, from material which we should See also:call not living . It abolished the conception of life as an entity above and beyond the See also:common properties of matter, and led to the conviction that the marvellous and exceptional qualities of that which we call " living " matter are nothing more nor less than an exceptionally complicated development of those chemical and physical properties which we recognize in a gradually ascending See also:scale of evolution in the See also:carbon compounds, containing See also:nitrogen as well as See also:oxygen, See also:sulphur and See also:hydrogen as constituent atoms of their enormous molecules . Thus mysticism was finally banished from the domain of biology, and zoology became one of the physical sciences—the science which seeks to arrange and discuss the phenomena of animal life and form, as the outcome of the operation of the laws of physics and chemistry .
A subdivision of zoology which was at one time in favour is simply into See also:morphology and physiology, the study of form
and structure on the one See also:hand, and the study of See also:scope
the activities and functions of the forms and structures or zoo-
on the other
.
But a logical See also:division like this is not 10g Y. necessarily conducive to the ascertainment and remembrance of the historical progress and present significance of the science
.
No such distinction of mental activities as that involved in the division of the study of animal life into morphology and physiology has ever really existed: the investigator of animal forms has never entirely ignored the functions of the forms studied by him, and the experimental inquirer into the functions and properties of animal tissues and organs has always taken very careful See also:account of the forms of those tissues and organs
.
A more instructive subdivision must be one which corresponds to the separate currents of thought and mental preoccupation which have been historically manifested in western Europe in the gradual evolution of what is to-day the great See also:river of zoological doctrine to which they have all been rendered contributory
.
It must recognize the following five branches of zoological study:
1
.
Morphography.—The work of the collector and systematist: exemplified by Linnaeus and his predecessors, by Cuvier, Agassiz, See also:Haeckel
.
2
.
Bionomics.—The See also:lore of the farmer, gardener, sportsman, fancier and field-naturalist, including thremmatology, or the science of breeding, and the allied teleology, or science of organic adaptations: exemplified by the See also:patriarch See also:Jacob, the poet See also:Virgil, See also:Sprengel, See also:Kirby and See also:Spence, See also:Wallace See also:anti Darwin
.
Zoo-See also:Dynamics, Zoo-Physics, Zoo-Chemistry.—The pursuit of the learned physician,—anatomy and physiology: exemplified by See also:Harvey, See also:Haller, Hunter, Johann See also: It is unnecessary to follow in this See also:article all these subjects, since they are for the most part treated under separate headings, not indeed under these names—which are too comprehensive for that purpose—but under those of the more specific questions which arise under each . Thus Bionomics is treated in such articles as EVOLUTION, HEREDITY, VARIATION, See also:MENDELISM, RE-See also:PRODUCTION, See also:SEX, &C.; Zoo-dynamics under MEDICINE, See also:SURGERY, PHYSIOLOGY, ANATOMY, See also:EMBRYOLOGY, and allied articles; Plasmology under See also:CYTOLOGY, PROTOPLASM, &c.; and Philosophical Zoology under numerous headings, EvoLUTIoN, BIOLOGY, &e . See also ZOOLOGICAL See also:DISTRIBUTION, PALAEONTOLOGY, OCEANOGRAPHY, See also:MICROTOMY, &C . It will be more appropriate here, without giving what would be a needless repetition of considerations, both historical and theoretical, which appear in other articles, to confine ourselves to two general questions, (I) the history of the various schemes of See also:classification, or Morphography, and (2) the consideration of the main tendencies is the study of zoology since Darwin . |
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